Leicester's Tom Youngs has been called into England's squad for the autumn internationals. Photograph: David Rogers/Getty Images

The uncapped forwards Mako Vunipola and Tom Youngs have been called into the England squad for the QBE autumn internationals.

Stuart Lancaster has made four injury-enforced changes to the elite squad he originally named in July for the Tests against Fiji, Australia, South Africa and New Zealand.

The Saracens prop Vunipola, whose father played at two World Cups for Tonga, and Leicester's hooker Young – the elder brother of the England scrum-half Ben Youngs – are covering for Alex Corbisiero and Rob Webber.

The Harlequins wing Ugo Monye has also been promoted to fill the back-three spot left vacant when Ben Foden damaged ankle ligaments playing for Northampton.

James Haskell, who returned to the England team for the drawn third Test against South Africa in Port Elizabeth on the summer tour, has been promoted from the Saxons to replace Tom Croft.