Mail Online's 'Paw-fect recipes for your pet's plate' was most shared national press story on Twitter in 2014

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A Mail Online story about a YouTube programme teaching dog owners how to prepare meals for their pets was the most shared national newspaper story on Twitter of 2014 according to new research.

A study by Searchmetrics found that links to The Guardian were tweeted more times than any other title, with an average of 392,358 tweets a week.

The Telegraph (307,690) and Mail Online (237,381) were the second and third most popular on the social networking site, and The Sun (3,531), Daily Star (6,556) and Times (8,772) were the least popular.

The study analysed the number of tweets per week generated by content on 11 newspaper websites – The Independent, Mirror, Financial Times, Daily Express and Daily Record in addition to those already mentioned.

Searchmetrics, a global search experience optimisation platform, also found the top ten tweeted stories of the year, with Mail Online’s “Paw-fect recipes for your pet's plate” shared 77,000 times.

Searchmetrics founder Marcus Tober said: “Interestingly our list of the most tweeted stories indicates it’s not always the biggest and most high profile news stories that get the most tweets. Often it’s the quirkier stories or those with a human-interest angle that catch people’s imagination and go viral.”