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The latter is commoner. In it the cell contains a pair of nuclei which fuse (see p. 95). Thereafter the basidium behaves as an asexual reproductive cell. The nucleus divides, usually into four, and each offspring-nucleus passes through an outgrowth (sterigma) into a rounded cell (basidiospore) which finally becomes broken off to reproduce the fungus. ZOOSPORANGIA These are sporangia within which the contents form z o o s p o r e s , often by division into 2, 4, 8, or more protoplasts. They are naked ASEXUAL REPRODUCTION 33 oval cells with flagella attached.

Longitudinal section of capsule of B 2 Funaria. 34 FORM AND STRUCTURE Asci, continued. forming around the protoplasm immediately surrounding each of the usually eight nuclei in the ascus. Besides this, these nuclei, which have been derived by free cell formation, are derived from those of rows of cells (ascogenous hyphae) which have grown out from a fertilized or unfertilized female organ. ). There is a good deal of sterile tissue ; in liverworts the spore-forming region is a rounded mass of spores, mixed with a proportion of elaters (long fibre-like cells which help to loosen the spores when the structure bursts open) ; in mosses the spore-forming region is a cylindrical or barrel-shaped mass, with a sterile columella down the middle, and a sort of lid, the operculum, a t the top ; the columella breaks down after the spores ripen, and the lid opens.

In adult plant cells there is often a single large central vacuole ± traversed by strands of protoplasm in which the nucleus may be suspended, or it may be lodged in protoplasm a t one side of the cell. The c e l l - s a p in such vacuoles although watery is not pure water, but has both colloidal and crystalloidal substances in it. Salts and soluble carbohydrates are included, especially sugar. Inulin which takes the place of starch in composite plants also occurs dissolved in the sap, although it is thrown out of solution in alcoholpreserved material in the form of globular crystalline masses.