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Don Diego Vega (Tyrone Power) is urgently called home by his father. To all outward appearances, he is the foppish son of wealthy ranchero and former Alcade Don Alejandro Vega (Montagu Love), having returned to California after his military education in Spain.

Don Diego is horrified at the way the common people are now mistreated by the corrupt Alcalde, Luis Quintero (J. Edward Bromberg), who had forced his father from the position of Alcalde. Don Diego adopts the guise of El Zorro ("The Fox"), a masked outlaw dressed entirely in black, who becomes the defender of the common people and a champion for justice.

In the meantime he romances the Alcalde's beautiful and innocent niece, Lolita (Linda Darnell), whom he grows to love. As part of his plan, Don Diego simultaneously flirts with the Alcalde's wife Inez (Gale Sondergaard), filling her head with tales of Madrid fashion and culture and raising her desire to move there with her corrupt husband, Luis.

In both his guises Don Diego must contend with the governor's ablest henchman, the malevolent Captain Esteban Pasquale (Basil Rathbone). He eventually dispatches the Captain in a fast-moving rapier duel-to-the-death, forcing a regime change, Don Diego's plan all along.

The Mark of Zorro is a sound remake of the lavish 1920 smash hitsilent film starring Douglas Fairbanks as Zorro and Noah Beery, Sr. as Sergeant Gonzales. That 1920 feature introduced Zorro's iconic all-black costume, subsequently incorporated into Johnston McCulley's later Zorro stories in his original fiction series upon which Fairbanks' film had been based. The 1920 movie was the first in a popular array of swashbuckler action features starring the acrobatic Fairbanks, who had previously appeared mainly in comedies. Clips from the film were incorporated into The Artist nine decades later.

In the DC Comics continuity it is established that The Mark of Zorro was the film that the young Bruce Wayne had seen with his parents at a movie theater, moments before they were killed in front of his eyes by an armed thug. Zorro is often portrayed as Bruce's childhood hero and an influence on his Batman persona. There are discrepancies regarding which version Bruce saw: The Dark Knight Returns claims it was the Tyrone Power version, whereas a story by Alan Grant claimed it to be the silent Douglas Fairbanks original. Bill Finger was himself inspired by Fairbanks' Zorro, including similarities in costumes, the "Bat Cave" and Zorro's cave, and unexpected secret identities, especially since the Batman character predates the Tyrone Power remake by a year. The posters for The Mark of Zorro and the 1981 film Excalibur were used for a scene in Batman v Superman: Dawn of Justice,[5]

In the animated series Justice League Unlimited, a flashback of the fateful night establishes that for DCAU continuity Bruce and his parents were attending The Mark of Zorro but does not indicate which version. In earlier episodes of Batman: The Animated Series, the fictional character Gray Ghost, a pulp fiction hero inspired by The Shadow, is the inspiration to young Bruce Wayne.