… live in burrows on sandy beaches above the strandline. Older individuals dig their burrows further from the sea. Burrows can be up to 1.3 metres (4 ft 3 in) deep, and can be closed off with sand during hot periods.

They can produce a variety of sounds, by striking the ground with the claw, by stridulation with the legs, and an incompletely explained “bubbling sound”. Males compete in a heavily ritualised manner which prevents the need for physical contact. O. quadrata is more active at night than in the day time, and is an omnivore, eating clams, insects, plant material, detritus and even other crabs..

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