While climate change has been recognised as threat to some of these sites, sea level rise has rarely been incorporated into management plans for their future preservation.

A recent UN report found that coastal regions are some of the most vulnerable to climate change, with millions likely to be forced from their homes in the coming decades as rising tides make flooding the norm and contaminate freshwater supplies.

Coastal protections are a feasible adaptation strategy for Unesco sites, and the authors of the new Nature Communications paper recommended that such measures should be integrated into the sites they identified.

One example of such a project already underway is the MOSE system in Venice, which will see the entire lagoon protected by submerged mobile barriers.

Crucially, the researchers noted that this system does not interfere with the appearance of Venice, or with the fragile ecosystem of the lagoon.

They said their findings should be used as a basis for local policy makers to make evidence-based decisions to protect their heritage for future generations.