This complex includes the species T. adamsi, T. latreillei, T. marojejy, T. nazgul, T. sabatra, and T. smaug. They are all characterized by the presence of very strongly developed longitudinal rugae on the forecoxae, and the absence of sculpture on the first gastral tergite (except in some specimens of T. manongarivo, which have costulate sculpture on the basal half of the first gastral tergite).