A spot opens up in Jess' downtown loft with the boys when Nick (Jake Johnson) contemplates moving in with his ex-girlfriend, Caroline (returning guest Mary Elizabeth Ellis). Lennon, 41, will play Nick's possible replacement, Neil.

In addition to playing Lieutenant Jim Dangle for six years on Reno 911!, Lennon is best known for the Night of the Museum films and How to Lose a Guy in 10 Days. He will also appear in the upcoming ABC comedy Apt. 23.