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Born Sydney, 14 April 1909. Died 26 March 1998. Educated University of Sydney (BScAgr 1932, DScAgr 1956). Field assistant, Griffith Research Station 1933, science master, Scotch College, Mitcham 1934, plant breeder, New South Wales Department of Agriculture 1935-36, research officer and botanist, Museum of Applied Science, Sydney 1937-47, Lecturer, then senior lecturer, then reader in Botany, University of Sydney 1948-63, professor of biology (genetics) 1963-74, Head, School of Biological Sciences 1966-67, honorary Associate, Universities of Sydney and New South Wales 1975-80. Fellow, Australian Academy of Science 1962, Clarke Medal, Royal Society of New South Wales 1968. President, Linnean Society of New South Wales 1958.