Lloyd M. Bertholf

Lloyd M. Bertholf, a MBL Corporation Life Member and former president of Illinois Wesleyan University died January 20, 2003 at age 103 in Bloomington, Illinois. A nationally regarded biologist, Bertholf was renowned for his research on the physiology and behavior of honeybees. A graduate of Southwestern College in Winfield, Kansas, he received his M.S. and Ph.D. degrees from Johns Hopkins University in Baltimore, where he studied zoology and chemistry. Under a Rockefeller fellowship, Bertholf conducted post-doctoral research into insect behavior in 1930-31 at the University of Munich in Germany, where he worked with famed scientist Karl von Frisch, who discovered how insects communicate.