Source: Social Facilitation, Affiliation, and Dominance in the Social Life of Spotted Hyenas. Stephen E. et al (http://www.fs.fed.us/psw/publications/zabel/zabel2.PDF)
Scent-Marking.
Hyenas scent-mark within their territories by communal defecating’ as well as by marking stalks of grass and other objects with “paste” extruded from their anal scent glands.
Spotted hyenas also display some rather unique characteristics. Adult females and their juvenile, or subadult, offspring dominate adult immigrant males in virtually all social interactions.4-6 In addition, y
h enas often spend their days alone at solitary dens, typically reassembling in the late afternoon and socializing at the communal
den before forming smaller hunting parties. For many hyenas there is a daily transition from a solitary existence to the intense, highly differentiated social interactions of life within the clan.