Background

Dr. Marvin H. Caruthers, Ph.D., co-founded miRagen Therapeutics, Inc. in 2006. Dr. Caruthers served as a Venture Partner and Consultant at Boulder Ventures Limited (BV) since its founding in 1995. From 1992 to 1995, he served as the Chairman of the Department of Chemistry and Biochemistry at the University of Colorado. In 1980, Dr. Caruthers and seven other research scientists founded AMGEN with three venture capital groups. He continues to serve as a Scientific Consultant ... to the AMGEN and serves a Member of the Scientific Advisory Board. Dr. Caruthers and Dr. Lee Hood co-founded Applied Biosystems, Inc., in 1981 to commercialize their discoveries in the chemical synthesis of nucleic acids and the sequencing of proteins. Dr. Caruthers is a distinguished Professor of Biochemistry and Bio-organic Chemistry at the University of Colorado since 1979. Dr. Caruthers is a renowned Research Scientist in the field of DNA Synthesis and an inventor and multiple patent holder of key enabling manufacturing techniques for the biotechnology industry. He served as a Director of Genomica Corp. since 1997 and has also been a Founding Investor at both the companies, together with BV General Partner Kyle Lefkoff. He has been a Director of ProFoldRx, Inc. since February 2006. He serves as Member of Scientific Advisory Board of AmideBio, LLC. Dr. Caruthers served as a Director of Dharmacon Inc., where he is also a Founding Investor. He serves as a Director of OXiGENE Inc. and miRagen Therapeutics, Inc. He has been a Director of BaroFold, Inc. since February 2006. He serves as a Member of Scientific Advisory Board at Nanosphere, Inc. He served as a Director of Array Biopharma Inc. from August 1998 to October 26, 2011. He served as a Member of Scientific Advisory Board at BiOptix Inc. Dr. Caruthers is an Editorial Board Member of the Journal of Biotechnology, the Journal of Protein Engineering, and the Journal of Molecular Recognition. He is a Member of the National Academy of Sciences and of the American Academy of Arts and Sciences. Dr. Caruthers was awarded the National Medal of Science for 2006, the nation’s highest distinction honoring scientific achievement. Dr. Caruthers, a Guggenheim Fellow, holds a Ph.D. in Biochemistry from Northwestern University and a B.S. in Chemistry from Iowa State University. He completed Post-Doctoral studies at MIT.

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