Lorne Gill said: “Choosing winners from the hundreds of images we receive is never easy, but the level of creativity and the diversity of photographs makes it such a pleasure.

“So many of the images are thought-provoking and inspiring.”

The winners received a range of prizes including trips to the Isle of May National Nature Reserve with Lorne Gill; memberships of the Scottish Seabird Centre and Scottish Wildlife Trust; vouchers for Northshots Photography and day-long wildlife trips courtesy of Isle of Mull Wildlife Expeditions.

An exhibition of the winning photographs will be on display at the Scottish Seabird Centre until later this month.