And they lived happily ever after...or so everyone was led to believe. Emma Swan knows how to take care of herself. She's a 28-year-old bail bondsperson who's been on her own ever since she was abandoned as a baby. But when Henry—the son she gave up years ago—finds her, everything changes. Henry is now 10 years old and in desperate need of Emma's help. He believes that Emma actually comes from an alternate world and is Snow White and Prince Charming's missing daughter. According to his book of fairytales, they sent her away to protect her from the Evil Queen's curse, which trapped the characters of fairytale world forever, frozen in time, and brought them into our modern world.

Not believing a word of Henry's story, Emma brings him back to Storybrooke, but finds herself drawn to this unusual boy and his strange New England town. Concerned for him, she decides to stay for a while, but soon comes to suspect that Storybrooke is more than it seems. It's a place where magic has been forgotten, but is still powerfully close...where fairytale characters are alive, even though they don't remember who they once were, and where the Evil Queen, known as Regina, is now Henry's foster mother. The epic battle for the future of all worlds is beginning, and for good to win, Emma will have to accept her destiny and fight like hell.

Wish Rumple's evil plan is revealed and Regina realizes the only hope to stop him is by turning Wish Henry from his path for vengeance. Meanwhile, Tilly and Margot try to get help in Storybrooke. With his hope of ever being reunited with Belle fading, Weaver struggles to find a way to defeat his evil alter ego; and when Rogers' life is threatened, he is faced with the ultimate sacrifice.

Alice returns home to her father telling her that everyone thought she was dead. Her imaginative stories about a talking White Rabbit and a mystical place called Wonderland end up getting her admitted into an asylum where Dr. Lydgate convinces her to have a procedure that will allow her to forget her pain. Right before the operation is set to take place she is rescued by the Knave of Hearts who takes her to the rabbit who informs her that her true love Cyrus is still alive. As the trio then jumps into the portal returning to Wonderland, Alice sets out on a quest to find him but the villainous Jafar and the Red Queen have other plans for her.

The all-powerful Jafar succeeds in making his father love him and forcing Anastasia into thinking she loves him. After Alice, Amara and Cyrus escape, Jafar imprisons the Jabberwocky and calls upon an army of dead soldiers to do his bidding. Meanwhile, Alice and the White Rabbit raise their own army to fight Jafar for the realm and a final confrontation between Alice and Jafar takes place at the Well of Wonders.

Emma Swan's life has been anything but a fairytale. A 28-year-old bail bondsperson, she's been taking care of herself since she was abandoned as a baby. But when Henry, the son she gave up for adoption 10 years ago, finds her, everything changes. Henry is desperate for his mom's help, and he thinks that Emma is actually the long-lost daughter of Snow White and Prince Charming.

Even stranger, Henry believes that Storybrooke, the sleepy New England town he calls home, is really part of a curse cast by the Evil Queen, freezing fairytale characters in the modern world with no memory of their fairytale selves. Of course, the seen-it-all Emma doesn't believe a word; but when she gets to Storybrooke, she can't help sensing that nothing is quite what it seems.

In Victorian England, the young and beautiful Alice tells a tale of a strange new land that exists on the other side of a rabbit hole. An invisible cat, a hookah smoking caterpillar and playing cards that talk are just some of the fantastic things she's seen during this impossible adventure. Surely this troubled girl must be insane and her doctors aim to cure her with a treatment that will make her forget everything. Alice seems ready to put it all behind her, especially the painful memory of the genie she fell in love with and lost forever — the handsome and mysterious Cyrus. But deep down Alice knows this world is real and just in the nick of time, the sardonic Knave of Hearts and the irrepressible White Rabbit arrive to save her from a doomed fate. Together, the trio will take a tumble down the rabbit hole to this Wonderland where nothing is impossible.

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