While training waterbomber captains for the upcoming fire-fighting season Chief Pilot Arnie makes a surprising decision. AJ becomes a DC-4 captain and ponders whether to accept a promotion. Meanwhile Joe and the whole McBryan family pile into the ol' DC-3 and head to the Super Bowl of aviation  AirVenture Air Show in Oshkosh Wisconsin.

Joe and Mikey head off to Coventry England to navigate the potential purchase of an Electra turboprop to add to Buffalo's fleet of fire-fighting tanker planes. The father-son duo makes a great team; hard-nosed Joe deals with the business end while Mikey bonds with the clients during a raucous visit to the local pub. Meanwhile new dad Rod leads a mission to Manitoba and misses his newborn daughter while away on the job.

When thunderstorms and a shaking engine threaten a DC-3 charter to Whitehorse, Joe pulls out all the stops to get the passengers to their destination. Patient C-46 co-pilot Scott Blue finally gets his crack at the DC-4, and the newest member of the McBryan dynasty, baby Emma Rae, makes her first visit to the Buffalo hangar.

Chief Pilot Arnie lands the DC-3 on a homemade sand strip at a remote lodge. Electra expert Chuck goes one-on-one with a stubborn power unit and Rod's wife Sasha goes into labour with the newest member of the McBryan family.