GemmaArtertonOnline has the first photo of Gemma Arterton in costume on the set of Louis Leterrier‘s Clash of the Titans. In the film, Arterton (Quantum of Solace) plays Io, Perseus’ mysterious spiritual guide throughout his journey. Clash is currently filming in Europe. Check out the previously released photos of Sam Worthington in action.

In Clash of the Titans, the ultimate struggle for power pits men against kings and kings against gods. But the war between the gods themselves could destroy the world. Born of a god but raised as a man, Perseus (Sam Worthington) is helpless to save his family from Hades (Ralph Fiennes), vengeful god of the underworld, Perseus volunteers to lead a dangerous mission to defeat Hades before he can seize power from Zeus (Liam Neeson) and unleash hell on earth. Leading a daring band of warriors, Perseus sets off on a perilous journey deep into forbidden worlds. Battling unholy demons and fearsome beasts, he will only survive if he can accept his power as a god, defy his fate and create his own destiny.

Rounding out the cast is Mads Mikkelsen (“Casino Royale”) as Draco, who takes up his sword to join Perseus’ quest; Jason Flemyng (The Curious Case of Benjamin Button) as Acrisius, a one-time king-turned-hideous beast; and Alexa Davalos (Defiance) as Andromeda, a princess doomed to lose her life if Perseus does not succeed.

Clash of the Titans is currently shooting, and is set for a March 26th 2010 release.