The panel of experts, who visited Guinea, said soldiers took part in mass murder and also carried out mass rape and sexual abuse of women at the protest, in the capital Conakry on 28 September.

The report, based on interviews with more than 600 people, said at least 109 girls and women were subjected to rape, sexual mutilation or kidnap for repeated rape.

Hundreds of people were also subjected to torture and abuse, it said.

Capt Camara had previously sought to distance himself from the atrocities by blaming unruly elements in the army.

But the report says: "The commission considers there are sufficient grounds for presuming direct criminal responsibility by President Moussa Dadis Camara."

Another passage states: "The commission recommends that the International Criminal Court be seized with respect to those persons on whom, according to this report's findings, weighs a strong suspicion of crimes against humanity."

Junta in turmoil

Critics of the junta had gathered in a sports stadium in the capital, Conakry, to protest at reports that Capt Camara intended to stand for president in an election planned for 2010.

But troops opened fire on the protesters in what human rights groups have described as a pre-planned massacre.

Guinea has been in turmoil since the military took over last December, but the shooting of Capt Camara by one of his soldiers on 3 December this year has thrown the country into even greater chaos.

Capt Camara was flown to Morocco for treatment after the shooting and has not yet returned - fuelling rumours that he was seriously injured.

The soldier implicated in the shooting, Lt Toumba Diakite, has been on the run ever since.

He has been accused of being in charge of some of the soldiers who opened fire on the protesters, and the UN panel recommended that he should also face charges.

Lt Toumba, in an interview with French media last week, said he had shot Capt Camara because he feared he would be blamed for the stadium killings.

A third member of the junta, the drug police chief Moussa Sakho Camara, was also named by the UN experts as bearing responsibility for the massacre.

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