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The England lock Simon Shaw has been ruled out of Saturday's Calcutta Cup clash against Scotland with a shoulder problem. The 36-year-old picked up the injury in the opening minutes of England's 20-16 defeat to Ireland at Twickenham on 27 February.

Martin Johnson confirmed this afternoon that the Wasps forward would not be available to play at Murrayfield and that the Stade Français lock Tom Palmer has been brought into the squad.

The England manager has called up five reinforcements in total, including Mike Tindall who returns to the senior squad after overcoming a hamstring injury. Tindall replaces the Leicester centre Dan Hipkiss, who is still struggling with an ankle problem.

Delon Armitage is said to be progressing well after injuring his ribs against Ireland but Johnson has turned to the Gloucester full-back Olly Morgan as cover. The Leeds flanker Hendre Fourie has received his first senior call-up after Wasps' Dan Ward-Smith suffered knee ligament damage at the weekend.

And the Northampton fly-half Shane Geraghty has returned to the senior fold as precautionary cover for Toby Flood, who received a knock to the head playing for Leicester against London Irish on Saturday. Flood's condition is currently being reviewed by the England medics.