But it raised even bigger questions about the origins of this ancient manuscript.

And there are now suggestions from the Middle East that the discovery could be even more spectacularly significant than had been initially realised.

There are claims that these could be fragments from the very first complete version of the Koran, commissioned by Abu Bakr, a companion of the Prophet Muhammad - and that it is "the most important discovery ever for the Muslim world".

This is a global jigsaw puzzle.

But some of the pieces have fallen into place.

It seems likely the fragments in Birmingham, at least 1,370 years old, were once held in Egypt's oldest mosque, the Mosque of Amr ibn al-As in Fustat.

Paris match

This is because academics are increasingly confident the Birmingham manuscript has an exact match in the National Library of France, the Bibliotheque Nationale de France.

Image copyrightThinkstockImage caption
The Mosque of Amr ibn al-As in Egypt, where Birmingham's Koran seems to have orginated

The library points to the expertise of Francois Deroche, historian of the Koran and academic at the College de France, and he confirms the pages in Paris are part of the same Koran as Birmingham's.

Alba Fedeli, the researcher who first identified the manuscript in Birmingham, is also sure it is the same as the fragments in Paris.

The significance is that the origin of the manuscript in Paris is known to have been the Mosque of Amr ibn al-As in Fustat.

'Spirited away'

The French part of this manuscript was brought to Europe by Asselin de Cherville, who served as a vice consul in Egypt when the country was under the control of Napoleon's armies in the early 19th Century.

Prof Deroche says Asselin de Cherville's widow seemed to have tried to sell this and other ancient Islamic manuscripts to the British Library in the 1820s, but they ended up in the national library in Paris, where they have remained ever since.

Image caption
1: In summer 2015 two leaves of an ancient Koran at the University of Birmingham were identified and dated as being much earlier than anyone had anticipated and among the oldest in the world. 2: The National Library of France, Paris has leaves from the same Koran, brought from Egypt by a vice consul under Napoleon. 3: The Mosque of Amr ibn al-As in Fustat, Egypt. The fragments of the Koran in Birmingham are believed to have come from this ancient mosque. 4: Alphonse Mingana was born near Zakho in modern-day Iraq in 1878. He brought the manuscript to Birmingham from the Middle East on a collecting trip in the 1920s funded by the Cadbury family.

But if some of this Koran went to Paris, what happened to the pages now in Birmingham?

Prof Deroche says later in the 19th Century manuscripts were transferred from the mosque in Fustat to the national library in Cairo.

Along the way, "some folios must have been spirited away" and entered the antiquities market.

These were presumably sold and re-sold, until in the 1920s they were acquired by Alphonse Mingana and brought to Birmingham.

Mingana was an Assyrian, from what is now modern-day Iraq, whose collecting trips to the Middle East were funded by the Cadbury family.

"Of course, no official traces of this episode were left, but it should explain how Mingana got some leaves from the Fustat trove," says Prof Deroche, who holds the legion of honour for his academic work.

And tantalisingly, he says other similar material, sold to western collectors could, still come to light.

Disputed date

But what remains much more contentious is the dating of the manuscript in Birmingham.

What was really startling about the Birmingham discovery was its early date, with radiocarbon testing putting it between 568 and 645.

The latest date in the range is 13 years after the death of the Prophet Muhammad in 632.

Image copyrightThinkstockImage caption
Manuscripts were brought to France by Napoleon's forces in Egypt

David Thomas, Birmingham University's professor of Christianity and Islam, explained how much this puts the manuscript into the earliest years of Islam: "The person who actually wrote it could well have known the Prophet Muhammad."

But the early date contradicts the findings of academics who have based their analysis on the style of the text.

Mustafa Shah, from the Islamic studies department at the School of Oriental and African Studies in London, says the "graphical evidence", such as how the verses are separated and the grammatical marks, show this is from a later date.

In this early form of Arabic, writing styles developed and grammatical rules changed, and Dr Shah says the Birmingham manuscript is simply inconsistent with such an early date.

Prof Deroche also says he has "reservations" about radiocarbon dating and there have been cases where manuscripts with known dates have been tested and the results have been incorrect.

'Confident' dates are accurate

But staff at Oxford University's Radiocarbon Accelerator Unit, which dated the parchment, are convinced their findings are correct, no matter how inconvenient.

Researcher David Chivall says the accuracy of dating has improved in recent years, with a much more reliable approach to removing contamination from samples.

Image caption
Jamal bin Huwaireb believes that this an even more remarkable find than has so far been realised

In the case of the Birmingham Koran, Mr Chivall says the latter half of the age range is more likely, but the overall range is accurate to a probability of 95%.

It is the same level of confidence given to the dating of the bones of Richard III, also tested at the Oxford laboratory.

"We're as confident as we can be that the dates are accurate."

And academic opinions can change. Dr Shah says until the 1990s the dominant academic view in the West was that there was no complete written version of the Koran until the 8th Century.

But researchers have since overturned this consensus, proving it "completely wrong" and providing more support for the traditional Muslim account of the history of the Koran.

The corresponding manuscript in Paris, which could help to settle the argument about dates, has not been radiocarbon tested.

The first Koran?

But if the dating of the Birmingham manuscript is correct what does it mean?

There are only two leaves in Birmingham, but Prof Thomas says the complete collection would have been about 200 separate leaves.

Image copyrightPAImage caption
The text of the Koran has remained unchanged since this very early version in Birmingham

"It would have been a monumental piece of work," he said.

And it raises questions about who would have commissioned the Koran and been able to mobilise the resources to produce it.

Jamal bin Huwareib, managing director of the Mohammed bin Rashid Al Maktoum Foundation, an education foundation set up by the ruler of the UAE, says the evidence points to an even more remarkable conclusion.

He believes the manuscript in Birmingham is part of the first comprehensive written version of the Koran assembled by Abu Bakr, the Muslim caliph who ruled between 632 and 634.

Image caption
Prof Thomas says the writer of this manuscript could have heard the Prophet Muhammad preach

"It's the most important discovery ever for the Muslim world," says Mr bin Huwareib, who has visited Birmingham to examine the manuscript.

"I believe this is the Koran of Abu Bakr."

He says the high quality of the hand writing and the parchment show this was a prestigious work created for someone important - and the radiocarbon dating shows it is from the earliest days of Islam.

"This version, this collection, this manuscript is the root of Islam, it's the root of the Koran," says Mr bin Huwareib.

"This will be a revolution in studying Islam."

This would be an unprecedented find. Prof Thomas says the dating fits this theory but "it's a very big leap indeed".

'Priceless manuscript'

There are other possibilities. The radiocarbon dating is based on the death of the animal whose skin was used for the parchment, not when the writing was completed, which means the manuscript could be a few years later than the age range ending in 645, with Prof Thomas suggesting possible dates of 650 to 655.

Media captionThe university's academics were "startled" by the results of radiocarbon dating