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"In the case of the larger wound, if the blade had penetrated 7cm into the brain, which we cannot determine from the bones, death would have been instantaneous."

Other wounds included slashes or stabs to the face and the side of the head. There was also evidence of "humiliation" injuries, including a pelvic wound likely to have been caused by an upward thrust of a weapon, through the buttock.

Richard III was portrayed as deformed by some Tudor historians and indeed the skeleton's spine is badly curved, a condition known as scoliosis.

However, there was no trace of a withered arm or other abnormalities described in the more extreme characterisations of the king.

Missing princes

Without the scoliosis, which experts believe developed during teenage years, he would have been about 5ft 8ins (1.7m) tall, but the curvature would have made him appear "considerably" shorter.

Dr Appleby said: "The analysis of the skeleton proved that it was an adult male but was an unusually slender, almost feminine, build for a man.

"Taken as a whole, the skeletal evidence provides a highly convincing case for identification as Richard III."

Richard was a royal prince until the death of his brother Edward IV in 1483. Appointed as protector of his nephew, Edward V, Richard instead assumed the reins of power.

Edward and his brother Richard, known as the Princes in the Tower, disappeared soon after. Rumours circulated they had been murdered on the orders of their uncle.

Challenged by Henry Tudor, Richard was killed at Bosworth in 1485 after only two years on the throne.

DNA trail

He was given a hurried burial beneath the church of Greyfriars in the centre of Leicester.

Mr Buckley said the grave was clumsily cut, with sloping sides and too short for the body, forcing the head forward.

University of Leicester findings

• Wealth of evidence, including radiocarbon dating, radiological evidence, DNA and bone analysis and archaeological results, confirms identity of last Plantagenet king who died over 500 years ago

• DNA from skeleton matches two of Richard III's maternal line relatives. Leicester genealogist verifies living relatives of Richard III's family

• Individual likely to have been killed by one of two fatal injuries to the skull - one possibly from a sword and one possibly from a halberd

• Ten wounds discovered on skeleton - Richard III killed by trauma to the back of the head. Part of the skull sliced off

• Radiocarbon dating reveals individual had a high protein diet - including significant amounts of seafood - meaning he was likely to be of high status

• Radiocarbon dating reveals individual died in the second half of the 15th or in the early 16th Century - consistent with Richard's death in 1485

• Skeleton reveals severe scoliosis - onset believed to have occurred at the time of puberty

• Although about 5ft 8in tall (1.7m), the condition meant King Richard III would have stood significantly shorter and his right shoulder may have been higher than the left

• Feet were truncated at an unknown point in the past, but a significant time after the burial

"There was no evidence of a coffin or shroud which would have left the bones in a more compact position.

"Unusually, the arms are crossed and this could be an indication the body was buried with the wrists still tied," he added.

Greyfriars church was demolished during the Reformation in the 16th Century and over the following centuries its exact location was forgotten.

However, a team of enthusiasts and historians managed to trace the likely area - and, crucially, after painstaking genealogical research, they found a 17th-generation descendant of Richard's sister with whose DNA they could compare any remains.

Joy Ibsen, from Canada, died several years ago but her son, Michael, who now works in London, provided a sample.

The researchers were fortunate as, while the DNA they were looking for was in all Joy Ibsen's offspring, it is only handed down through the female line and her only daughter has no children. The line was about to stop.

Tomb plans

But the University of Leicester's experts had other problems.

Dr Turi King, project geneticist, said there had been concern DNA in the bones would be too degraded: "The question was could we get a sample of DNA to work with, and I am extremely pleased to tell you that we could."

She added: "There is a DNA match between the maternal DNA of the descendants of the family of Richard III and the skeletal remains we found at the Greyfriars dig.

"In short, the DNA evidence points to these being the remains of Richard III."

In August 2012, an excavation began in a city council car park - the only open space remaining in the likely area - which quickly identified buildings connected to the church.

Richard's villainous reputation owes much to the way he was characterised by Shakespeare

The bones were found in the first days of the dig and were eventually excavated under forensic conditions.

Details of the reburial ceremony have yet to be released, but Philippa Langley from the Richard III Society said plans for a tomb were well advanced.

She said of the discovery of Richard's skeleton: "I'm totally thrilled, I'm overwhelmed to be honest, it's been a long hard journey. I mean today as we stand it's been nearly four years.

"It's the culmination of a lot of hard work. I think, as someone said to me earlier, it's just the end of the beginning.

"We're going to completely reassess Richard III, we're going to completely look at all the sources again, and hopefully there's going to be a new beginning for Richard as well."

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