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Dani Moreno (Katusha) won Flèche Wallonne with a well-timed attack on the final climb up the Mur de Huy to win ahead of Sergio Henao (Sky) and Carlos Betancur (AG2R-La Mondiale).

Moreno bided his time when Betancur charged off the front with 700 metres of the climb still to go. World champion and pre-race favourite Philippe Gilbert (BMC) took up responsibility for the chase halfway up the climb but although his forcing managed to shed Peter Sagan (Cannondale), he played his hand too early.

Inside the final 200 metres, Moreno came off the fading Gilbert’s shoulder and then scorched past Betancur to take the win. Henao came through for second place, while Betancur narrowly held off the rapidly-closing Dan Martin (Garmin-Sharp) to finish third.

A disappointed Gilbert almost ground to a halt in the closing 150 metres, while neither Joaquim Rodriguez (Katusha) nor Alejandro Valverde (Movistar) made an impression on the steep slopes of the Mur de Huy as the day belonged to their fellow countryman Moreno.

It was a race that was always going to come down to the Mur de Huy, despite the peloton at one point being in three separate groups split apart by considerable aggression from a number of teams following the first ascent of the taxing 1300 metre climb.

Vos led out the sprint with 100 metres remaining, dropping her pursuants' wheels, and the victory never looked in doubt as the prolific Dutchwoman had enough time to sit up and celebrate in emphatic style, as the other two battled it out for the minor placings on the line.

The in-form Italian Longo Borghini was left frustrated on the ground while she waited for confirmation of her final result to go along with her gut instinct, but with that came lots of smiles.

"We were really close but I had a feeling that I was number two," the 21-year-old told Cyclingnews. "I was just waiting for the jury."

Longo Borghini's result on Wednesday isjust the latest in what has been an impressive spring, claiming top-10 performances at Le Samyn des Dames, Omloop van het Hageland, the Tour of Flanders and well as her victory at the Trofeo Alfredo Binda - Comune di Cittiglio World Cup race.

"I'm really happy for this race and for the whole spring," Longo Borghini said. "My team did an amazing job. I really have to thank all my teammates and of course the staff.

"The last climb of the Mur was really hard. Vos went from around 100 metres and I tried to follow but in the end it was a sprint of two between me and Moolman."

Moolman's result was one for the history books, the first South African woman to podium in a World Cup race as well as her best result at the top level.

"I must say it's quite overwhelming," the 27-year-old told Cyclingnews. "I'm standing here today as a very proud South African and very proud African ambassador.

"I think I went a little too early," she said of the sprint to the line. "I got a little too excited; I could hear my manager on the radio saying ‘Use the steep part, use the steep part.' I think I tried to lunge for the second place but it was just so close."

The peloton was intact on the first climb of the day despite an early attack from Dani King (Wiggle Honda). Aleksandra Burchenkova (RusVelo) was next to attack, earning 20 seconds on the bunch, but she too was brought back.

Just under 50km of racing was done when the peloton shattered into action with eight riders gaining 42 seconds on the Côte de Bohissau. The peloton in two distinct groups, with the chase well and truly on, the eight leaders were caught and it was gruppo compatto once more.

The first ascent of the Mur de Huy at 65km saw splits emerge once more, with a large chunk of the peloton abandoning as the selection continued. As the riders re-fuelled, the race was back together and the finale was on the way was nearing.

Late attacks by Alena Amialiusik (BePink) and Katie Colclough (Specialized – lululemon) proved fruitless, never able to gain more than 30 seconds of clear air.

Former mountain biker Tatiana Guderzo (MCipollini Giordana) was prepared to give it one more crack however in the final 10km but the wider roads proved to be her downfall and her 10 second advantage was easily swallowed up, setting up Vos for another victory.