In paleogeography, Gondwana/ɡɒndˈwɑːnə/,[1][2] also Gondwanaland, is the name given to the more southerly of two supercontinents (the other being Laurasia) that were part of the Pangaea supercontinent that existed from approximately 510 to 180 million years ago (Mya). Gondwana formed prior to Pangaea, then became part of Pangaea, and finally broke up after the breakup of Pangaea. Gondwana is believed to have sutured between about 570 and 510 Mya, thus joining East Gondwana to West Gondwana.[3] It separated from Laurasia 200-180 Mya (the mid-Mesozoicera) during the breakup of Pangaea, drifting farther south after the split.[4]

The adjective "Gondwanan" is in common use in biogeography when referring to patterns of distribution of living organisms, typically when the organisms are restricted to two or more of the now-discontinuous regions that were once part of Gondwana, including the Antarctic flora. For example, the Proteaceae family of plants known only from southern South America, South Africa, Australia, and New Zealand is considered to have a "Gondwanan distribution". This pattern is often considered to indicate an archaic, or relict, lineage.

Other blocks which helped to form Argentina and some surrounding regions, including a piece transferred from Laurentia when the west edge of Gondwana scraped against southeast Laurentia in the Ordovician.[6][7] This is the Famatinian block (named after Famatina in northwest Argentina) and it formerly continued the line of the Appalachians southwards.[8]

Reconstruction showing final stages of assembly of Gondwana, 550 Mya

One of the major sites of Gondwanan amalgamation was the East African Orogeny (Stern, 1994), where these[which?] two major orogenies are superimposed. The East African Orogeny at about 650–630 Mya affected a large part of Arabia, north-eastern Africa, East Africa, and Madagascar. Collins and Windley (2002) propose that in this orogeny, Azania collided with the Congo–Tanzania–Bangweulu Block.[9]

The later Malagasy orogeny at about 550–515 Mya affected Madagascar, eastern East Africa and southern India. In it, Neoproterozoic India collided with the already combined Azania and Congo–Tanzania–Bangweulu Block, suturing along the Mozambique Belt.[10]

At the same time, in the Kuunga Orogeny Neoproterozoic India collided with the Australia/Mawson continent.

Other large continental masses, including the core cratons of North America (the Canadian Shield or Laurentia), Europe (Baltica), and Siberia, were added over time to form the supercontinent Pangaea by Permian time. When Pangaea broke up (mostly during the Jurassic), two large masses, Gondwana and Laurasia, were formed.

The reformed Gondwanan continent was not precisely the same as that which had existed before Pangaea formed; for example, most of Florida and southern Georgia and Alabama is underlain by rocks that were originally part of Gondwana, but that were left attached to North America when Pangaea broke apart.[11]

During the late Paleozoic, Gondwana extended from a point at or near the South Pole to near the Equator. Across much of Gondwana, the climate was mild. During the Mesozoic, the world was on average considerably warmer than it is today. Gondwana was then host to a huge variety of flora and fauna for many millions of years. The laurel forest of Australia, New Caledonia, and New Zealand have a number of other related species of the laurissilva de Valdivia, through the connection of the Antarctic flora as gymnosperms and deciduous angiosperm Nothofagus. Corynocarpus laevigatus is called the bay of New Zealand, Laurelia novae-zelandiae belongs to the same genus Laurelia. The sempervirens treeniaouli grows in Australia, New Caledonia, and New Zealand.

New Caledonia and New Zealand ecoregions became separated from Australia by continental drift 85 million years ago. The islands still retain plants that originated in Gondwana and spread to the Southern Hemisphere continents later. However, strong evidence exists of glaciation during the Carboniferous to Permian time, especially in South Africa.

The Nothofagus plant genus illustrates Gondwanan distribution, having descended from the supercontinent and existing in current day Australia, New Zealand, New Caledonia, and the Southern Cone. Fossils have also recently been found in Antarctica.[12]

Gondwana began to break up in the early Jurassic (about 184 Mya) accompanied by massive eruptions of basalt lava, as East Gondwana, comprising Antarctica, Madagascar, India, and Australia, began to separate from Africa. South America began to drift slowly westward from Africa as the South Atlantic Ocean opened, beginning about 130 Mya during the EarlyCretaceous, and resulting in open marine conditions by 110 Mya. East Gondwana then began to separate about 120 Mya when India began to move northward.

As the age of mammals commenced, the continent of Australia-New Guinea began gradually to separate and move north (55 Mya), rotating about its axis to begin with, and thus retaining some connection with the remainder of Gondwana for about 10 million years.

About 45 Mya, the Indian Plate collided with Asia, buckling the crust and forming the Himalayas. At about the same time, the southernmost part of Australia (modern Tasmania) finally separated from Antarctica, letting ocean currents flow between the two continents for the first time. Antarctica became cooler and Australia became drier because ocean currents circling Antarctica were no longer directed around northern Australia into the subtropics.

The separation of South America from West Antarctica some time during the Oligocene, perhaps 30 Mya, also caused climate changes. Immediately before this separation, South America and East Antarctica were not connected directly. However, the many microplates of the Antarctic Peninsula remained near southern South America, acting as "stepping stones" and allowing continued biological interchange and stopped oceanic current circulation. When the Drake Passage opened, there was no longer a barrier to force the cold waters of the Southern Ocean not to be exchanged with warmer tropical water. Instead, a cold circumpolar current developed and Antarctica became what it is today: a frigid continent that locks up much of the world's fresh water as ice. Sea temperatures dropped by almost 10°C, and the global climate became much colder.

By about 15 Mya, the collision between New Guinea (on the leading edge of the Australian Plate) and the southwestern part of the Pacific Plate pushed up the New Guinea Highlands, causing a rain shadow effect which drastically changed weather patterns in Australia, drying it out.

^H.M. Li and Z.K. Zhou (2007) Fossil nothofagaceous leaves from the Eocene of western Antarctica and their bearing on the origin, dispersal and systematics of Nothofagus. Science in China. 50(10): 1525-1535.