GspA is homologous to exeA of A. hydrophila, which is required for protein export, specifically for the export and assembly of secretin (ExeD/GspD) into the outer membrane. GspA has an N-terminal predicted cytoplasmic domain with an ATP/GTP binding motif, a single predicted transmembrane segment (239-261), followed by a predicted periplasmic domain with a predicted C-terminal peptidoglycan-binding motif (407-467; Pfam PG_binding_1). GspA may interact with GspB. A divergon is a regulon comprised of two adjacent divergent operons or genes. Overexpression causes a decrease in the amount of biofilm formed and decreases motility.