Thermal models of subduction zones often base their slab-wedge geometry from seismicity at mantle depths and, consequently, cannot be used to evaluate the relationship between seismicity and structure. Here, high-resolution seismic observations from the recent Broadband Experiment Across the Alaska Range (BEAAR) constrain, in a rare instance, the subducting slab geometry and mantle wedge temperature independent of seismicity. Receiver functions reveal that the subducting crust descends less steeply than the Wadati-Beinoff Zone. Attenuation tomography of the mantle wedge reveals a high Q and presumably cold region where the slab is less than 80 km deep. To understand these two observations, we generate thermal models that use the improved wedge geometry from receiver functions and that incorporate temperature- and strain-rate-dependent olivine rheology. These calculations show that seismicity within the subducting crust falls in a narrow belt of pressure-temperature conditions, illuminating an effective Clapeyron slope of 0.1 K/N4Pa at temperatures of 450-750 degrees C. These conditions typify the breakdown of high-pressure hydrous minerals such as lawsonite and suggest that a single set of dehydration reactions may trigger intermediate-depth seismicity. The models also require that the upper, cold nose of the mantle wedge be isolated from the main flow in the mantle wedge in order to sustain the cold temperatures inferred from the Q tomography. Possibly, sufficient mechanical decoupling occurs at the top of the downgoing slab along a localized shear zone to 80 km depth, considerably deeper than inferred from thrust zone seismicity. (c) 2005 Elsevier B.V. All rights reserved.