Several thousand years have passed since the demon warrior Sparda raised his sword against the unholy world to save the human race. Victorious and assured that evil forces had been confined, Sparda joined mankind and fathered Dante, a half human-half demon son infused with the power of a supernatural lineage. However, Dante was not the only son Sparda had sired. As devoted as Dante was to defeating the forces of evil, his twin brother, Virgil, was consumed by power and upheld the legacy of his sinister ancestors. One moonlit night, a mysterious tower materialized beside Dante's "Devil May Cry" investigative agency. On the rooftop stood his brother along with his inhuman minions goading Dante into a confrontation.