"Female differs from modestus, panamensis, and harterti by the more rufous general coloration, and browner, less blackish head. Male browner above than modestus, and more cinnamon buff below" (Blake 1977: 32)

Occurs on the Pacific coast of Panama, including Isla del Rey in the Pearl Islands.

panamensis Carriker 1910

"Female browner below than male, and both sexes much darker than poliocephalus; crown and hindneck blacker; foreneck and upper breast darker gray. Similar to modestus in general appearance, but darker, more reddish brown above; male darker, less ochraceous below; undersurface of female less grayish than in modestus" (Blake 1977: 32).

Occurs on the Pacific slope in Panama from the Río Majé (Panamá) east to Darién; and on the Caribbean slope of Panama from western Colón east.

harterti Brabourne and Chubb 1914

"A transitional form connecting the browner or duskier Central American races with the more rufous forms of upper Amazonia. Nearest caucae, but female more grayish brown, less rufescent above, and much less rufescent below; lower foreneck and breast more decidedly grayish, less tawny. Male differs from caucae in darker general coloration and less ochraceous underparts; from caquetae in grayer, less brownish dorsum, and relatively obscure barring on the flanks and tibiae" (Blake 1977: 32).

Occurs on the Pacific coast of Colombia, Ecuador, and extreme northwestern Peru (Tumbes), north to the Gulf of Urabá.

caucae Chapman 1912

"Female darker than mustelinus, and with a sooty pileum; differs from the nominate race only in having the chest strongly tinged with dusky gray. Male similar to mustelinus below and slightly paler than soui; uppersurface browner, less rufescent than mustelinus" (Blake 1977: 33).

Occurs in northern Colombia in the middle Magdalena valley and in the drainages of the Cauca and Sinú rivers.

mustelinus Bangs 1905

"Male closely similar to the nominate race, but less rufescent above, and more buffy, less tawny below. Female usually paler above than soui, and with brownish instead of sooty pileum" (Blake 1977: 33).

Occurs in northeastern Colombia (Santa Marta region south to Boyacá), and northwestern Venezuela in Zulia, Mérida and northern Táchira.

andrei Brabourne and Chubb 1914

"Closely similar to the nominate race, but upperparts, sex for sex, darker and less rufescent. Female virtually indistinguishable from soui below, but ventral coloration of male less bright, dull tawny olive, rather than clay color" (Blake 1977: 33).

Occurs from southeastern Colombia east to the Guianas, and Brazil north of the Amazon.

caquetae Chapman 1915

"A transitional form between mustelinus and soui, and the dark extreme nigriceps. Male not certainly separable from soui; differs from mustelinus in darker and browner dorsum, and darker ochraceous, less olivaceous underparts. Female above and below also darker and browner than mustelinus, pileum sooty, and chest with a dusky band; slightly darker and browner than both soui and caucae, and chest distinctly dusky" (Blake 1977: 34).