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co-adaptation

The year 1870 is aptly summarised by the brief entry Darwin made in his journal: ‘The whole of the year at work on the Descent of Man & Selection in relation to Sex’. Descent was the culmination of over three decades of observations and reflections on human ancestry, including the origin of language, mind, morals, and religious temperament. The year was otherwise coloured by controversies, including vigorous objections to the application of natural selection to humans from Wallace and St George Jackson Mivart.

Bates’s letter is that of a true thinker. Asks to see JDH’s to Bates. Point raised in it is most difficult. "There is one clear line of distinction; – when many parts of structure as in woodpecker show distinct adaptation to external bodies, it is preposterous to attribute them to effect of climate etc. – but when a single point, alone, as a hooked seed, it is conceivable that it may thus have arisen." His study of orchids shows nearly all parts of the flower co-adapted for fertilisation by insects and therefore the result of natural selection.Mormodes ignea"is a prodigy of adaptation".

Outlines his theory on the origin of existing orders of leaf arrangement. Believes spiral and whorled orders have evolved from a primitive distichous arrangement. These arrangements permit a compact bud form of small surface area that can withstand external changes in temperature, and in particular can tolerate frost.