Footnotes

Author contributions: S.W.B. and G.I.L. designed research; S.W.B., R.E.P., and G.I.L. performed research; S.W.B. and G.I.L. contributed new reagents/analytic tools; S.W.B. and G.I.L. analyzed data; and S.W.B. and G.I.L. wrote the paper.

The authors declare no conflict of interest.

This article is a PNAS Direct Submission.

Data deposition: The short-read sequencing data reported in this paper have been deposited in the NCBI BioProject database (accession no. PRJNA388205).