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I had one of the Flatline drops when I had a Flatline tank. It was really cool for quickly disconnecting that tank.

From that picture, though, I can't understand how those work. The two pieces that look like they'd fit together, is there a tiny lip on the sides to keep them stuck together or something?

It's kind of tough to see in the picture, but there are two horizontal dowel pins that rotate into the slots milled into the side of the receiver. The two pieces must be perpendicular before being rotated. The tiny lever on the end moves an internal cam which pushes the ball bearings upward onto the hardened steel inserts of the other piece.