But restrictions have been lifted after a "crucial final test" came back all clear.

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Staff are "looking to the future with a renewed sense of certainty" after the disease affected breeding and animal movement programmes for endangered animals such as red panda, golden lion tamarin and the Asiatic lion, the zoo said.

Dr Amy Plowman, director of living collections, said: "We’ve been able to make a few selected animal moves recently with the help of government vets at either end, thanks to an agreement with the Animal and Plant Health Agency.

"We hope now that we can accelerate the process, moving youngsters on, pairing up single animals and thinking about our conservation breeding work."