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Stefan Themerson was born in Płock in what was then part of the Russian Empire on 25 January 1910 and died in London on 6 September 1988. His father, Mieczysław Themerson, was a physician, social reformer and aspiring writer (some of his work was published) of Jewish descent. His mother was Ludwika Smulewicz. During the First World War Dr. Themerson served as a medical officer in the Tsar's army and the family lived in Riga, St. Petersburg and Wielkie Luki. In 1918 they returned to Płock, in an independent Poland, where Stefan attended the Władysław Jagiełło Gymnasium. During this time he first showed an interest in photography and also built a radio receiver. In 1928 Themerson went to Warsaw University to study physics. After a year, he transferred to the Warsaw Polytechnic to take up architecture but actually spent most of his time working at photography, collage and film-making. His first published piece of writing was also in 1928. He never formally left his studies but gradually withdrew to follow his other interests. Themerson met the brilliant art student Franciszka Weinles, in 1929. She graduated with distinction in 1931 and they married the same year in Warsaw.

During these years the Themersons lived and worked in Warsaw. Stefan contributed articles to various periodicals and prose and verse to school textbooks and wrote at least ten books for children which Franciszka illustrated. Pan Tom Buduje Dom, Mr Rouse Builds His House is still in print in Poland. Stefan also experimented with photograms and the two of them made five short experimental films AptekaThe Pharmacy (1930), Europa (1931–1932), Drobiazg Melodyjny, Musical Moment (1933), Zwarcie, Short Circuit (1935) and Przygoda Człowieka Poczciwego, The Adventures of a Good Citizen (1937). These were shown with other experimental films of the time. All except Przygoda Człowieka Poczciwego, which remained in Warsaw, were lost in Paris in the Second World War, but the script for Europa, based on a poem by Anatol Stern was later published by the Themersons Gaberbocchus Press, illustrated by surviving stills from the film and Apteka was remade from descriptions of it when it first appeared, stills and storyboards. In 1935, with other young filmmakers, they founded a cooperative, S.A.F, Spółdzielnia Autorów Filmowych.

In 1936 and 1937 the Themersons visited Paris, then the centre of the world for avant-garde art, and London, meeting Moholy-Nagy and other experimental artists and arranged first screenings of French and English on their return to Warsaw. They also founded a review Film Artistique, Stefan as editor, Franciszka as artistic editor (the journal of the film-makers' co-operative). Two issues were published, one on the French avant-garde, one on recent English film. A third issue, on the Polish Avant-garde, was planned but remained unpublished when, in the winter of 1937-38 they moved to Paris: "I just knew I had to be in Paris." Themerson said. Here they found a circle of artists and writers, many Polish, to live and move in. Themerson wrote for various Polish publications in Paris and Franciszka started to paint. They planned to stay there, but - as for most of their generation - the War changed their lives. With the outbreak of war in 1939, both of the Themersons enlisted, Stefan joined the Polish army forming in France after the German and Soviet invasions and division of Poland, Franciszka was seconded to the Polish government-in-exile as a cartographer, first in France then from 1940 in London.

In 1940 Themerson volunteered for a Polish infantry regiment in France, just in time for the German invasion and the Allies' collapse. His memory was of marching day and night in the summer heat to St-Nazaire. There, in June, the regiment was disbanded, the officers abandoning their men and the men dispersing where they could. Themerson travelled inside France, first returning (on foot) to occupied Paris, then on to Toulouse, where, through the Polish Red Cross, he re-established contact with Franciszka. She had been working for the Polish Government in Exile as a cartographer in Paris and Normandy and had subsequently escaped to London on a troopship from Bayonne. Meanwhile, he spent time in refugee camps, worked as a farm labourer, and spent over a year in a Polish Red Cross hostel, the Hôtel de la Poste in Voiron. Here he began writing Professor Mmaa's Lecture in Polish and wrote the long poem Croquis dans les Ténèbres, Sketches in Darkness. Much of their sporadic correspondence from the years 1940-42 concerned attempts to engineer Stefan's escape from France. Finally, towards the end of 1942 Themerson got across France via Marseille and Spain to Lisbon where he was flown to Britain by the R.A.F., rejoining his wife and re-enlisting in the Polish army. He spent time with the army in Scotland, where he finished Professor Mmaa, and then was sent to join the film unit of the Polish Ministry of Information and Documentation in London. There he and Franciszka made two further short films, Calling Mr Smith, an account of Nazi atrocities in Poland and The Eye and the Ear, inspired by four Julian Tuwim songs set to music by Szymanowski. In 1944 at the PEN club meeting to celebrate the three hundredth anniversary of John Milton's Areopagitica, Themerson met Kurt Schwitters, who became a close friend until his death. At about the same time he met others who remained close, including Jankel Adler, Julian Trevelyan and Anthony Froshaug. Also in 1944 the Themersons moved to Maida Vale, where they lived for the rest of their lives. A close neighbour was the experimental poet and publisher Bob Cobbing with whom the Themersons kept close relations.

Stefan and Franciszka Themerson published many of their own works through their highly original Gaberbocchus Press from 1948 to 1979, most were designed by Franciszka, and many included her illustrations. They also worked with the translators Barbara Wright and Stanley Chapman. Their list of some 70 titles included works by Guillaume ApollinaireJankel Adler and Kurt Schwitters, the first English translation of Alfred Jarry's Ubu Roi, Raymond Queneau's Exercises in Style and The Good Citizen's Alphabet by Bertrand Russell. The latter wrote an encouraging preface to Professor Mmaa’s Lecture. Stefan was the inventor of Semantic Poetry, (see his novel 'Bayamus' 1945, and 'On Semantic Poetry' 1975), and he wrote an opera, 'St. Francis and the Wolf of Gubbio' (1972). In 1981, Themerson delivered the annual Huizinga Lecture in Leiden in the Netherlands, under the title: The Chair of Decency. He died in London a few weeks after the death of his wife, in 1988.