Bowser is at it again with his son and the Koopalings, but this time things are much different: he didn't kidnap Princess Peach! Rather, Bowser went and kidnapped a number of Sprixies, a species of fairy creatures that are greenish in appearance. Mario and friends pursue Bowser through a strange pipe to the Sprixie Kingdom to rescue the Sprixies and once again stop Bowser's plot!