"Some of the more unusual tracks include 'tippy-toe' traces - this is where fully buoyed dinosaurs made deep, near vertical scratch marks with their toes as they propelled themselves through the water.

"It's difficult to see how tracks such as these could have been made by running or walking animals. If that was the case we would expect to see a much flatter impression of the foot preserved in the sediment."

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Mr Romilio said the swimming dinosaur tracks at Lark Quarry belonged to small, two-legged herbivorous dinosaurs known as ornithopods.

"These were not large dinosaurs. Some of the smaller ones were no larger than chickens, while some of the wading animals were as big as emus," he said.