The Imperial Star Destroyer Infiltrator was commisioned in 22 BBY during the Clone Wars. It saw limited service defending the Core Worlds before the end of the conflict.

After the conclusion of the Clone Wars, the Infiltrator was put into active duty suppressing uprisings and pursuing rememants of the Seperatist Military. In 18 BBY, the Infiltrator was assigned to the Maridis Sector as part of the Sector Group under the command of Admiral Terrel Harkness. Command of the vessel was transferred to the Admiral's daughter, High Inquisitor Syanne Harkness, shortly after that.

The Infiltrator has seen action against ships of the Alliance to Restore the Republic, Antrixian Resistance forces, and Dominion Imperium ships.