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Before the end of the Time War, this Dalek fell through time and landed on the Ascension Islands circa 1962. Insane and screaming, it passed through several private collections in the 20th and 21st centuries. By 2012, it was in the possession of billionaire Henry van Statten, who kept it in the Vault. Having no other name for it, van Statten called it a "Metaltron". The "Metaltron" was kept locked in the Cage, restrained by tensile steel chains attached to stabilising pillars. He instructed another of his staff, Simmons, to make it talk. The last person to touch it without any gloves "burst into flames", at least according to what Simmons said. Although tortured, all it did was scream. With the intention of reaching other Daleks, it sent out a distress signal which was detected by the Ninth Doctor and Rose Tyler. Initially unaware of the source of the distress signal, the Doctor came to face-to-face with it. (TV: Dalek) Having thought all of the Daleks destroyed (TV: The Day of the Doctor), he was shocked to find one alive. It moved its gun to exterminate the Doctor, but had insufficient power to do so. After a hostile and bitter discussion between them about the end of the Time War, the Doctor attempted to destroy the Dalek, pulling a lever which sent electricity through its casing. He was stopped by van Statten, who ordered two guards to "get him out", pulling the Doctor out the room. Simmons managed to turn the electricity off before the Dalek was destroyed. Henry van Statten was pleased to know what the "Metaltron" was really called.

Upon meeting Rose, who had not witnessed the exchange between the Dalek and the Doctor, it seemed to be a harmless victim. Rose, in an attempt to comfort it, touched the right part of the Dalek's dome (to the left of the eye stalk, from Rose's perspective). As she did so, it absorbed artron energy and DNA from her. It used this to regenerate itself and escape the Vault. It then downloaded all the data available on the Internet and killed Simmons, Bywater, De Maggio, and numerous others. It pursued both Rose and Adam Mitchell through the GeoComTex facility and killed 200 of van Statten's security.

However, the "Metaltron" found itself ultimately unable to kill Rose or van Statten. The human DNA that it had absorbed from Rose had brought with it emotions. Rose took the Dalek to the roof, where it blasted a hole in the ceiling and opened its battle armour to feel the sunlight. The Doctor then appeared, armed with an alien energy weapon, with the intent to kill the Dalek. Upon seeing it feel, the Doctor lowered the weapon at Rose's request.

Considering all the new emotions to be "sickness", the "Metaltron" asked Rose to order it to self-destruct, preferring death to a life with emotions. She refused at first, but eventually gave the order, and it destroyed itself. (TV: Dalek)

While it was a prisoner, the "Metaltron" refused to speak when Henry van Statten's staff were torturing it, although it did scream. It was enraged by the appearance of the Doctor and tried to exterminate him, but couldn't because its gun had been disabled. The Dalek demanded to be ordered what to do, ironically using orders itself in an attempt to receive them. However, it also begged for pity when the Doctor attempted to destroy it, despite the Doctor pointing out "you never did".

After absorbing Rose Tyler's DNA, the Dalek started feeling human emotions for the first time. Although it ruthlessly gunned down van Statten's staff, it was unable to kill Rose or van Statten. The Dalek then stated that it wanted freedom. As its personality had become more merciful than before, it hated itself, feeling that the emotions Rose had given it were making it weak. It eventually pled with Rose to order it to self-destruct, and she did so. (TV: Dalek)

This Dalek also acts as the player character in the online game The Last Dalek. It presents an alternate version of the events of Dalek. In the game, the Dalek fights through the Vault, faces van Statten's forces and eventually the Doctor, who is exterminated by the Dalek. The game is completed when the Dalek finds and destroys the Doctor's TARDIS.

In The Day of the Doctor, as Gallifrey vanishes and the Dalek fleet is destroyed, one Dalek seems to survive as it spins off from the explosion. Although this would be consistent with the general backstory of the "Metaltron", there is no clear indication that it is the same Dalek.