The death of Optimus Prime is a seminal moment for those who grew up with the toys, comics, and cartoon series. With the Matrix of leadership passed on to a brand-new character in Ultra Magnus (Robert Stack), the Autobots attempt to survive unaware that an even greater threat in the world-eating Unicron (Orson Welles) is looming. Taking the battered Megatron (Frank Welker) and remaking him into his lieutenant Galvatron (Leonard Nimoy) along with new soldiers, Unicron hopes to crush the Autobots.