Biology
Glossary
(e.g. epibenthic)

Usually inhabits lower and middle reaches of rivers and lakes. A sea-run form is found in the eastern coast of Hokkaido. Young feed mainly on aquatic insects while fishes more than 30 cm are completely piscivorous. Spawns in spring in river branches, where the female lays 2,000-10,000 eggs on sandy or gravelly bottom.