Patrick Norton wrote-
> These were all flyers, so quill knobs should be expected. What I am
> anticipating is evidence of quill knobs on feathered theropods outside
> Avialae--those non-flying maniraptorans such as Protarchaeopteryx or its
> relatives.
Archaeopteryx and Caudipteryx definitely lack ulnar quill knobs. The
specimens have been examined thoroughly and are well preserved. Only
Rahonavis among non-pygostylian theropods has quill knobs to my knowledge.
Mickey Mortimer