It is typical for wolves to put their rear paws into (or close to) the prints of their forepaws, and this happens crossed-over, so they put the right rear paw into the print of the left forepaw, and that's what happened right at the moment of the capture.

Great shot! I love how he looks like he's going to jump on you in any second. I just can't figure out, what's with his rare legs? Are they crossed? Looks like they either crossed, or his right rare leg (looking from wolf's perspective) is seriosly crooked. Anyway, the photo is great, I've been looking for something like this, to give my tattoo artist an idea, what I want on me