The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

This data file contains the data on weights and lengths from retrospective growth analysis of different stem age classes of Betula nana ramets from the LTER Nutrient and Warming manipulations in tussock tundra at Toolik Lake.

The Biocomplexity Station was established in 2005 to measure landscape-level carbon, water and energy balances at Imnavait Creek, Alaska. The station is now contributing valuable data to the AON project that was established at two nearby stations. These will form part of a network of observatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

Numbers of Eriophorum vaginatum inflorescences, both unclipped and clipped by small mammals, were counted in experimental plots. The plots are setup in moist acidic tussock tundra near Toolik Field Station, Alaska ((8 degrees 37' 27" N, 149 degrees 36' 27"W) and include fenced exclosures in both fertilized and unfertilized tundra.

Thaw depth was measured using a steel probe in the Imnavait Creek watershed, near Toolik Lake, Alaska. The thaw grid includes measurements made from the valley bottom (on both sides of the stream), up the hillslope to the hilltop (watershed boundary). The thaw grid is near Imnavait water tracks 7 and 8, and measurements have been made from the 2003 season until present. Two surveys are conducted each summer, on 2 July and on 11 August (plus or minus 1-2 days on either side of those dates).

Starting in 1990 thaw depth has been measured in a small watershed of tussock tundra every July 2nd and August 11th. The UTM stake grid (stakes are in a grid at every 100 meters) was used to locate the points . All measured depths are in cm. Measurements are made within a 20 cm radius of each location point. All the measurements are then averaged into a single value for the day.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

In contribution to the Arctic Observing Network (AON), the researchers have established two observatories of landscape-level carbon, water and energy balances at Imnaviat Creek, Alaska and at Pleistocene Park near Cherskii, Russia. These will form part of a network of obervatories with Abisko (Sweden), Zackenburg (Greenland) and a location in the Canadian High Arctic which will provide further data points as part of the International Polar Year.

The Multiple Element Limitation (MEL) model is used to simulate the recovery of Alaskan arctic tussock tundra to thermal erosion features (TEFs) caused by permafrost thaw and mass wasting. TEFs could be significant to regional carbon (C) and nutrient budgets because permafrost soils contain large stocks of soil organic matter (SOM) and TEFs are expected to become more frequent as climate warms. These simulations deal only with recovery following TEF stabilization and do not address initial losses of C and nutrients during TEF formation.

A study investigating the mechanisms that control long-term response of tussock tundra to fire and to increases in air temperature, CO2, nitrogen deposition and phosphorus weathering. The MBL MEL was used to simulate the recovery of three types of tussock tundra, unburned, moderately burned, and severely burned in response to changes in climate and nutrient additions. The simulations indicate that the recovery of nutrients lost during wildfire is difficult under a warming climate because warming increases nutrient cycles and subsequently leaching within the ecosystem.

This material is based upon work supported by the National Science Foundation under Grants #DEB-1637459, 1026843, 9810222, 9211775, 8702328; #OPP-9911278, 9911681, 9732281, 9615411, 9615563, 9615942, 9615949, 9400722, 9415411, 9318529; #BSR 9019055, 8806635, 8507493. Any opinions, findings, conclusions, or recommendations expressed in the material are those of the author(s) and do not necessarily reflect the views of the National Science Foundation.