"Hitler himself is believed to have had two forms of genital abnormality: an undescended testicle and a rare condition called penile hypospadias in which the urethra opens on the under side of the penis," Mr Mayo and Ms Craigie wrote.

The discovery could offer some explanation as to why Hitler was allegedly afraid of being seen naked and the cause of his famed fits of rage.

It also likely to add fuel to the debate on Hitler's sex life - or lack thereof - which is fiercely contested by historians.

In his biography of Hitler, the British historian Ian Kershaw said the Austrian-born Nazi leader was repelled by sexual activity of any kind as he feared catching an infection.

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