Department of Cellular and Molecular Pharmacology, University of California, San Francisco, San Francisco, CA 94158, USAHoward Hughes Medical Institute, University of California, San Francisco, San Francisco, CA 94158, USACalifornia Institute for Quantitative Biomedical Research, University of California, San Francisco, San Francisco, CA 94158, USA

Department of Urology, University of California, San Francisco, San Francisco, CA 94158, USAHelen Diller Family Comprehensive Cancer Center, University of California, San Francisco, San Francisco, CA 94158, USA

Department of Cellular and Molecular Pharmacology, University of California, San Francisco, San Francisco, CA 94158, USAHoward Hughes Medical Institute, University of California, San Francisco, San Francisco, CA 94158, USACalifornia Institute for Quantitative Biomedical Research, University of California, San Francisco, San Francisco, CA 94158, USA