Wilson et al. 2014. A female elephant browsing on Phyllanthus embellica. Invasive weeds such as Lantana camara have now covered significant parts of the reserve leading to a decline in native forage species. (Photo credit: Gaius Wilson)

Wilson et al. 2014. In this study a trunksful of grass was defined as the number of trunksful of grass or browse that the focal elephant gathered with its trunk and put into its mouth. Here, a herd of elephants is seen feeding on grass in a Lantana camara-free area of Mudumalai Tiger Reserve. (Photo credit: Gaius Wilson).