Acclaimed UC Irvine geneticist Francisco J. Ayala has resigned after a university investigation found he sexually harassed four faculty members and graduate students, the university announced Thursday.

Ayala, a former Dominican priest who has won the prestigious Templeton Prize for championing both science and faith, resigned effective July 1 and will abstain from future university activities, UCI officials said. He could not immediately be reached for comment.

In 2011, Ayala donated $10 million to the School of Biological Sciences, which then bore his name. It was the largest gift from a faculty member at the time.

The university said Ayala’s name has been removed from that school, and is also being removed from its central science library, graduate fellowships, scholar programs and endowed chairs. The biology school will now be known as the UCI School of Biological Sciences.