KHARTOUM, Sudan 
The Sudanese army said Sunday that it killed the leader of the main Darfur rebel group in fighting earlier this week, touting his death as a key victory against a powerful rebel force that once threatened Sudan's capital.

Khalil Ibrahim led the Darfur-based Justice and Equality Movement, or JEM, the most organized and effective military force in Darfur, the western region torn by conflict since 2003. The group did not join a peace deal signed last year in Doha, Qatar between other Darfur rebel groups and Sudanese President Omar al-Bashir's government in Khartoum.

The military said he was wounded Thursday during a military offensive in North Kordofan state, which borders Darfur. The government said he died of his wounds Saturday and that rebels quickly buried him. The government did not say how it confirmed his death.

JEM representatives could not immediately be reached for comment. If Ibrahim's death is confirmed by the group, it would be a serious blow to JEM, which has on several occasions threatened to bring down al-Bashir's regime in Khartoum by advancing toward the capital.

Sudanese Information Minister Abdullah Massar said Ibrahim's death sends a message to rebel groups "to listen to the voice of wisdom and join the peace process."

"Our doors are open and the Doha agreement is open," Massar said Sunday.

Darfur has been in turmoil since 2003, when ethnic African rebels accusing the Arab-dominated Sudanese government of discrimination took up arms against it. The Khartoum government is accused of retaliating by unleashing Arab militias on civilians - a charge the government denies.

The conflict has tapered off since 2009, but the U.N. estimates 300,000 people died and 2.7 million have been displaced. The International Criminal Court in the Hague has issued an international arrest warrant for al-Bashir for his alleged role in crimes against humanity in Darfur.

Sudan's Information Minister told reporters Sunday that 30 people were killed with Ibrahim in the clashes. An army statement carried by Sudan's official SUNA news agency said Ibrahim and several associates were killed in Wad Banda, about 440 miles (700 kilometers) west of Khartoum in the North Kordofan region, which borders Darfur.

"The army cut all paths for the group as it was retreating and trying to cross into South Sudan to reorganize it forces," said Sudanese army spokesman Col. Sawarmy Khaled. "Our gallant armed forces were able to kill rebel Khalil Ibrahim along with some of his associates."

South Sudan seceded from the north in July to become the world's newest nation.

Fighting in Wad Banda had flared up in the past few days. On Saturday, the Sudanese army said Darfur rebels attacked three locations in neighboring North Kordofan, killing an unspecified number of civilians.

JEM has not claimed responsibility for the attack. However, in a dramatic push in 2008, hundreds of JEM fighters drove through the remote western region and attacked Khartoum's western outskirts. More recently, JEM spread its armed presence from Darfur into North Kordofan.

A security officer with the United Nations Mission in Darfur told The Associated Press that JEM mostly operates out of North Kordofan state and no longer has an official presence in Darfur. He said UNAMID officials have not confirmed Ibrahim's death. He spoke on condition of anonymity because he was not authorized to speak to reporters.

This year, UNAMID had nearly 26,000 troops and police stationed in Darfur, making it the largest UN peacekeeping operation.

Salma Turabi, daughter of longtime opposition leader Hassan Turabi, said she was at a gathering of mourners at Ibrahim's home in Khartoum when police fired tear gas to disperse the crowd.

Sudan state television broadcast images of Khartoum residents outside military headquarters, waving Sudanese flags and celebrating the news of his death.

Ibrahim is believed to have recently returned to Sudan after years in exile in neighboring Libya, where he enjoyed the patronage of Moammar Gadhafi before the longtime Libyan dictator was ousted and killed in October. Sudanese officials said the JEM has been trying to regroup since losing Gadhafi's reported support for the rebel movement.

Ibrahim once served in al-Bashir's government before joining the rebellion. He refused to join a peace agreement signed in Qatar and backed by the African Union that was meant to provide a basis for a cease-fire, power sharing, equal distribution of wealth and compensation for displaced people.

"He completely refused to come to the negotiating table, he never joined the peace talks," Ismail el-Haj Musa, Sudanese deputy speaker of the council of states, told the pan-Arab Al-Jazeera TV on Sunday. "He committed acts against the state."

On Sunday, Ibrahim Mahmoud Hamad, Sudan's interior minister, described the death of Ibrahim "as a message to all rebels and those carrying arms, calling on them to the negotiating table to resolve issues and reach a peaceful resolution for the best interest of the country," according the semiofficial Sudan Media Center.