Cor funnelform or salverform, with or without fornices; anthers included or partly exserted; gynobase low-pyramidal or flat or depressed; nutlets smooth to pitted or wrinkled, basally attached, the large scar often surrounded by a sharp rim, sometimes only one nutlet maturing; annual to more often perennial herbs, seldom pungently hairy, with mostly yellow or white or greenish-white fls in modified leafy-bracteate cymes, or solitary in or near the upper axils, often heterostylic; fruiting pedicels mostly erect or ascending. (Buglossoides) 75, widespread, mostly temperate and mountainous regions.