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FIGURE 1 Temporal or age polyethism in the paper wasp Polybia aequatorialis. Data were collected on 130 individually marked, known-age workers. Two measures of worker activity at three task sets are plotted against worker age (solid lines: mean rate of task performance; dashed lines: number of workers performing the task). Note the typical centripetal developmental sequence: in-nest tasks (mostly nest cleaning) are followed by building on the nest exterior, and later by foraging (leaving the nest and returning with food and building materials).