1Division of Mucosal Immunology, Department of Microbiology and Immunology, The Institute of Medical Science, University of Tokyo, Tokyo, Japan.
2Department of Pediatrics, Osaka City University Graduate School of Medicine, Osaka, Japan.
3Division of Clinical Immunology, Karolinska Institutet at Karolinska University Hospital Huddinge, Stockholm, Sweden.
4Crop Development Division, NARO Agriculture Research Center, Niigata, Japan.
5Medical Proteomics Laboratory, The Institute of Medical Science, University of Tokyo, Tokyo, Japan.
6Enteric Virus Unit, Virus Reference Department, Health Protection Agency, London, United Kingdom.
7Unilever Research and Development Vlaardingen BV, Vlaardingen, Netherlands.
8Institute of Infection and Global Health, University of Liverpool, Liverpool, United Kingdom.
9International Research and Development Center for Mucosal Vaccine, The Institute of Medical Science, University of Tokyo, Tokyo, Japan.

Department of Microbiology and Immunology, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo 108-8639, Japan; Division of Bacteriology, Department of Infectious Diseases Control, International Research Center for Infectious Diseases, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo 108-8639, Japan.

Division of Mucosal Immunology, Department of Microbiology and Immunology, The Institute of Medical Science, The University of Tokyo, Tokyo 108-8639, Japan
Department of Medical Genome Science, Graduate School of Frontier Science, The University of Tokyo, Chiba 277-8562, Japan
International Research and Development Center for Mucosal Vaccines, The Institute of Medical Science, The University of Tokyo, Tokyo 108-8639, Japan
Laboratory of Vaccine Materials, National Institute of Biomedical Innovation, Osaka 567-0085, Japan
Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Tokyo 102-0075, Japan
Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, Japan
Department of Microbiology and Immunology, Columbia University Medical Center, New York, New York 10032, USA
Laboratory of Host Defense, WPI Immunology Frontier Research Center, Osaka University, Osaka 565-0871, Japan
Laboratory of Immune Regulation, Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Osaka 565-0871, Japan

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