American Association for the
Advancement of Science (Fellow)
Entomological Society of America (Emeritus)
International Society of Hymenopterists
International Union for the
Study of Social Insects North Carolina
Entomological Society
Sigma Xi
Society for the Study of Evolution

biogeography; evolution of sex;
sexual selection; the writings of Charles Darwin; species and
speciation;
life histories and their evolution; tropical insect ecology;
polyphenisms in behavior and ecology

University
of Kansas
Non-majors' course:Insects
and Man
Graduate course:Social
Insects

University
of Arizona,
University of Arkansas, University
of California – Los Angeles,
University
of Colorado,
Universidad de Costa Rica, University of Georgia (x2), Harvard University,
University of Illinois
(x2),IndianaStateUniversity, Iowa State
University, University
of Kansas
(x4), KansasStateUniversity,
University of Kentucky,
University
of Michigan,
University of Missouri - Columbia (x4),
North CarolinaStateUniversity
(x3), University of North Carolina, The OhioStateUniversity,
Saint Louis University (x3)