He said the pair left their vehicle and went in different directions in search of crocodiles.

A search and rescue operation was launched after Mr Van Zyl’s dogs returned to their camp without him.

The hunter’s footprints were traced to river bank alongside his discarded rucksack. He was married, had two children and took foreign clients on hunting trips.

Mr Louwrens said staff from the Heritage Protection Group – an organisation which he heads and helps police fight crime in South Africa – informally helped the Zimbabwean authorities conduct the search.

“Permission was given for three Nile crocodiles in the area to be shot, and one of them contained Mr Van Zyl’s remains,” he said. “Subsequent DNA tests have proved the remains to be those of Mr Van Zyl.”

At least four fatal attacks by crocodiles have been reported in Zimbabwe this year.

One conservation group meanwhile has condemned the circumstances of his “senseless” death.

“[He] shouldn’t have been hunting in the first place. Animals in the wild… are wild! They are living, thinking beings with instincts for survival,” a statement by One Green Planet said.

Nile crocodiles

Image copyrightScience Photo Library

Image caption
The Nile crocodile is on of Africa’s most feared predators

Predators of Africa’s rivers and lakes, they lurk almost totally submerged in the water as they lie in wait for passing prey which is then dragged into the water and drowned

Renowned for their long powerful jaws, few creatures escape from their clutches – not even large buffalo and wildebeest

Their large bodies and long tails hide quick reflexes and fast bursts of speed

They are extremely attentive and protective parents, with a surprisingly delicate touch – their nests and young hatchlings are highly vulnerable to predators