This is an Open Access Article. It is published by Elsevier under the Creative Commons Attribution 4.0 Unported Licence (CC BY). Full details of this licence are available at: http://creativecommons.org/licenses/by/4.0/

Sponsor:

This article arises, in part, from research co-funded by the Natural Environment Research Council (NERC grant number: NE/P012264/1 and NE/K00722X/1) and the Environment Agency (Project number:
SC 130021).