Lyoto landed 13 strikes over the course of the whole fight. Lyoto literally had to sprint at JBJ to hit him, meanwhile JBJ attacked safely from range. Gus isn't a sprinter, and even then JBJ has already seen that trick so I am sure he would be ready.

And yet Machida looked better and most had him winning the first round with his striking...

Jones didn't have success in that fight until he took Lyoto down. It ended pretty quickly after that though.

Imo Jones's biggest weakness is definitely his striking. His unnatural range and agility makes it hard for other fighters to get into range and then when they do get close enough they are in his takedown range which is even more dangerous for them.