The game is set during a series of unnatural disasters across Planet Earth due to unexplained appearances of large crystals, where the Vortaak alien race are invading once again. The game features over 20 Kaiju and Mechas from all three Godzilla eras; Shōwa, Heisei and Millennium, as well as two newly created Toho approved creations; Krystalak and Obsidius.