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The time is 1973. The Cold War of the mid-20th Century continues to damage international relations. Britain's Secret Intelligence Service (SIS), a.k.a. MI6 and code-named the Circus, is striving to keep pace with other countries' espionage efforts and to keep the U.K. secure. The head of the Circus, known as Control (John Hurt), personally sends dedicated operative Jim Prideaux (Mark Strong) into Hungary. But Jim's mission goes bloodily awry, and Control is forced out of the Circus — as is his top lieutenant, George Smiley (Gary Oldman), a career spy with razor-sharp senses.

Estranged from his absent wife Ann, Smiley is soon called in to see undersecretary Oliver Lacon (Simon McBurney); he is to be rehired in secret at the government's behest, as there is a gnawing fear that the Circus has long been compromised by a double agent, or mole, working for the Soviets and jeopardizing England. Supported by younger agent Peter Guillam (Benedict Cumberbatch), Smiley parses Circus activities past and present. In trying to track and identify the mole, Smiley is haunted by his decades-earlier interaction with the shadowy Russian spy master Karla.