In one of the least verbose of his chapters, Greebledotus gives us his version of the story of The Five Lions (breaks in the original papyrus are indicated by ... )

"...of which the users were said to gain wondrous powers. Originally designed to guard an alien castle in a faraway dimension, the five Lions were not cognizant and had to be piloted by very human looking creatures.
These machines...indeed...mighty opponents. All together, the five guardians would combine to create a powerful and gigantic robot to dispatch the mightiest of foes. Now, at some time in the past...spread to the farthest reaches of the...only...Then, by a confederacy of coincidences, they ended up in the hands of the dreaded forever young magicians...(long break in the text)
...one time. Imbued (read trapped) with their new spirits, they acquired the moniker 'The Five Lions'. Now, even more powerful than ever, the Five Lions broke the magical bonds that required them to heed the words of the Sorcerors."

Greebledotus goes on to explain each of the Lions and it's powers.

" ...of which the first, after...once is now known as Tamajishi. The original pilot of this lion was called 'Pidge', and his story is mostly unknown. At that time...until...then called Jamjamros...but eventually manifested sentience as the Jade Lion.
This Lion makes up the left arm of the giant robot, and ,embodies the qualities of defense, healing, and combat flexibility. This Lion has powerful defensive capabilities. In it's giant form this Lion holds the shield. It can ward off attacks on itself and the area around it, including magic. The Lion can heal all types of damage, including mechanical, and can even resurrect a dead helot or ten.
Little has been written about the offensive options for the Jade Lion, but even in it's oldest role, it was always noted for it's speed and initiative. "

In deference to the Ragnarok theme i would now like to exhibit what is currently known of the Blue Lion.
First up is Brikcydides, who in his own spoiled rich-kid, my family owns a good mine way, does offer an alternate history of the Blue lion's early history.

"The original pilot of the Blue Lion was known as Sven, and was caught outside his mechanical lion unawares by a group of magic users. Horribly injured, Sven retired to the hospital planet of Arus. He was later replaced by his younger, more handsome brother, Ryan."

Though different, Greebledotus has a somewhat modified version:

"The Blue Lion, now called Fireslaght Kottr, was originally piloted by a being known as High White Platinum. This pilot was exceptionally skilled, good looking, and strong. Unfortunately, his luck was to be caught by a dreaded witch outside his craft. Armed with only a light pistol, he managed to shoot out the right eye of his adversary before he was turned into a Holiday mince meat pie. He was later consumed by the witch's demonic children in a devilish ritual.

His brother, whose name is unknown to us, set out to avenge his kin's terrible demise. He travelled to a planet called Altea, with a large retinue of space marines, samurai warriors, and armored robots. Ultimately victorious, the brother then took control of the lion, although now cursed by those dreaded prevaricators of black magic. He was relegated to winter battles and Northern climate operations. The nature of the curse was as follows; since the Blue Lion's attacks were electricity based, the sorcerors tied this to generated heat. This caused the Blue Lion to overheat in almost all conditions, save the coldest. It was after this that the Blue Lion eventually ended up abandoned and lost to the glaciers of time.

When all of the Lions were reunited on the planet Arus, Fireslaght Kottr was the last one brought back. He had apparently ended up on the planet of Gaia, where he was taken care of for many years by the people known as Sami. It was they who appended his current name (Greebledotus references a separate name used only by the Sami).

After reunification, the curse was lifted, and Fireslaght Kottr could finally bear his tremendous offensive capabilities. The Blue Lion has the least amount of combat support and helots, mainly because his followers are relegated to their home planet and are not a space faring people.
Of course, he hardly needs it. His resistance to magic is highest amongst the lions, as is his resistance to cold and electricity based attacks. He can generate and control a powerful electromagnetic field, and uses it to fly, create a defensive block, or cast out like a giant mace.

Fireslaght Kottr can not only invoke lightning anywhere on the battlefield, but can also breathe lightning over a large number of opponents with a chainlike trajectory, leaping through entire battalions. The Blue Lion also has an impressive array of death beams, bladed melee weapons, and traditional guns, cannons, missiles, and rockets. His presence on the battlefield gives all of his teammates an offensive boost irregardless of proximity.

Fireslaght also has a somewhat aloof attitude towards combat, with a reluctance toward his potential damage perhaps limited by his violent past. If a battle goes on for to long, the Blue Lion has been known to withdraw from boredom and return to Gaia, to replenish his soul. "

Greebledotus had less to say about the Red Lion:
"The Red Lion makes up the right arm of the final assembled form of the giant guardian. Originally, it was piloted by a humanoid from the planet of Altea known as Brave Black Iron, whose hot-headedness and combat efficiency were renowned in his time.
In addition to an energy bite, shoulder lasers, and thermite dorsal rockets, the Red Lion also has a variety of heat and fire-based attacks. The massive wave of heat from his mouth is known as the 'pyroclastic flow', and can engulf whole squads of warriors.
In his release from the magical seal that had been placed on all five lions, he became known as Popoki Pele, and resides in a ring of volcanic activity on the planet of Gaia."

Tilophon, who actually fought alongside the Five Lions, wrote at least one detailed treatise on the Red Lion. A remnant of which Grebbledotus was aware of, but apparently not in possession of. It is part of a larger lost work, called 'Golionica':

"Known as Red Lion, he is driven by Isamu. He is autonomous, unless joined as the right arm of the Universal Guardian. Isamu is the highest rated marksman in the entire defense force. As a pilot his performance is rated as 'exemplary '. Only outmatched by his fellow pilots, he qualified to drive the Red Lion after the entire defense force was screened for the best pilots.
In his Guardian form, he holds the Sword of the Ten Kings, a reference to the lineage of royalty on the planet of Altea. Aggressive and brazen, Isamu is almost always the first pilot to enter into combat. His main combat tactic is to directly engage the most powerful opponent he can discern; and then perform a flanking maneuver, trying to blast large numbers of troops with his powerful rocket and laser batteries.
Isamu loves to race the Red Lion, and often be can be found cruising the beaches and oceans of Altea. Only outpaced by the Green Lion, Isamu works closely with the team of engineers and scientists responsible for the upkeep and advancement of the Lion technologies.
Isamu also works closely with the self defense forces to improve his fencing skills and empty hand fighting. Along with Seido, he has developed a set of special techniques, unique to the Guardian final form, that deal with close combat against giant foes. "