An independent vet inspected the puppies and determined that six were in need of treatment for sickness, and all the animals are now in the RSPCA's care.

A man was arrested on suspicion of breaching a ban on keeping animals, while a woman was arrested on suspicion of aiding and abetting the male to breach that ban.

Four dead puppies were found during the raids, which are part of Operation Pagan

Mobile phones and computers were seized from the addresses, and the pair were later bailed.

Chief inspector Ian Briggs, from the RSPCA's special operations unit, said: "There are thousands of dogs in rescue centres desperately looking for new homes, but many people continue to fuel the trade in imported puppies by buying from rogue sellers who simply see the animals as money makers.

"At this time of year especially, we want to hammer home the message that people should never buy a dog as a Christmas present.

"Having a pet is a major responsibility, and not something that should be done on a whim as a gift for someone.

"Never let your heart rule your head and buy a puppy because you feel sorry for it. If you have any reservations at all, you should always walk away."

Sergeant Mike Cooper, from Greater Manchester Police, said: "We have worked extremely closely with our colleagues at the RSPCA in the planning and execution of this operation.

"Two people have been arrested and are now on police bail pending further enquiries. We witnesses some shocking scenes yesterday and we will do everything we can to assist the RSPCA in any future prosecution or operation.

"People should only buy pets through the proper channels after considered thought and I would like to echo the RSPCA's sentiments that they should not be given as gifts this Christmas."

Illegal Animal Trade: 11 Of The Dumbest Animal Smugglers (PHOTOS)

PHOTOS: 11 Dumbest Animal Smugglers -- Lizards In Underwear, Endangered Species On eBay And More

PHOTOS: 11 Dumbest Animal Smugglers -- Lizards In Underwear, Endangered Species On eBay And More

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Songbird Socks

46-year-old Sony Dong was arrested at Los Angeles International Airport in 2009 for attempting to smuggle 15 songbirds from Vietnam. Who would have thought that feathers and bird poop on his feet would give him away? Dong used a custom cloth contraption that secured the birds to his calves via buttons, according to the AP.