In manipulation, parasites and memetic symbionts manipulate their hosts for their own ends. Since symbiont reproduction requires contact between hosts, making the hosts be nice to other hosts helps the symbionts to reproduce.

In overgeneralisation, resource-limited cognition results in organisms being nice to one another - because that is what usually pays.

As with other group selection theorists, Wilson may also be over-estimating the actual incidence of altruistic acts. For example, many forms of altruism act as a form of virtue signalling - and show how nice you are.