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The D558 program was conceived as a joint NACA/U.S. Navy research program for transonic and supersonic flight. As originally envisioned, there would be three phases to the D558 program: a jet-powered airplane, a mixed rocket/jet-powered configuration, and a design and mockup of a combat aircraft.[1] A contract for design and construction of six D558-1 aircraft for the first phase was issued on 22 June 1945. The original plan had been for six aircraft with a mixture of nose and side inlets and varying wing airfoil sections. That plan was quickly reduced to three aircraft of a single configuration with a nose inlet. Plans for the second phase with mixed rocket/jet propulsion were also dropped. Instead, a new aircraft, the D558-2, was designed with mixed rocket and jet propulsion for supersonic flight.

Construction of the first 558-1 began in 1946 and was completed in January 1947. The fuselage used magnesium alloys extensively, while the wings were fabricated from more conventional aluminum alloys. The airframe was designed to withstand unusually high loads of up to 18 times gravity due to the uncertainties of transonic flight. The forward fuselage was designed so that it, including the cockpit, could be jettisoned from the aircraft in an emergency.[2] The aircraft was configured to carry more than 500 lb of test equipment, including sensors (primarily strain gauges and accelerometers) in 400 locations throughout the aircraft. One wing was pierced by 400 small holes to enable aerodynamic pressure data to be collected.[3]

The Skystreaks were powered by one Allison J-35-A-11 engine (developed by General Electric as the TG-180) — one of the first axial-flow turbojets of American origin — and carried 230 US gallons (871 l) of aviation fuel (kerosene).

All the Skystreaks were initially painted scarlet, which led to the nickname "crimson test tube." NACA later had the color of the Skystreaks changed to white to improve optical tracking and photography. The first of three D-558-1 Skystreaks, BuNo 37970, made its maiden flight on 14 April 1947, at Muroc Army Air Field (later named Edwards AFB). Less than 4 months later, on August 20, this aircraft with Commander Turner Caldwell, USN, set a new world speed record of 640.744 miles per hour (1,031.178 km/h) flying D-558-1 #1, the very first air speed record that exceeded the unofficial mark of 1,004 km/h (623.8 mph) set by an example of the World War II-era German Me 163A Komet rocket fighter prototype, itself set in very early October 1941. The D-558-1 #1 Skystreak's record lasted 5 days, and was broken by then-Lieutenant Colonel Marion Carl, USMC, going 10 miles per hour (16 km/h) faster in D-558-1 #2, BuNo 37971. This aircraft was delivered to the NACA Muroc Flight Test Unit in April 1949 after 101 flights had been completed by the Navy, Air Force, and Douglas. This aircraft was never flown by the NACA. The D-558-1 #1 is located at the National Naval Aviation Museum at Naval Air Station Pensacola, Florida.

Following 27 flights by the Navy and Douglas the second D-558-1 aircraft was delivered to the NACA in November 1947. The D-558-1 #2 underwent extensive instrumentation by the NACA Muroc instrumentation section. The number 2 Skystreak made a total of 19 flights with the NACA before it crashed on takeoff due to compressor disintegration on May 3, 1948, killing NACA pilot Howard C. Lilly. The third D-558-I, BuNo 37972, aircraft was delivered to the NACA Muroc Flight Test Unit in 1949 after three Douglas test pilots and Howard Lilly had flown it. The number three aircraft took over the planned flight program of the D-558-1 #2. From the first flight in 1949 through 1953 the third Skystreak was flown in an intensive flight-research program by seven NACA test pilots, with a great deal of useful data collected on high-subsonic handling. The D-558-1 #3 made a total of 78 research flights with the NACA before being retired on June 10, 1953. The third Skystreak is on display at Carolinas Aviation Museum located at the Charlotte-Douglas International Airport (CLT) in Charlotte, North Carolina.

The Skystreak reached Mach 0.99 in level flight, but only flew supersonic in a dive.[3] In the public mind, much of the research performed by the D-558-1 Skystreaks was quickly overshadowed by Chuck Yeager and the supersonic Bell X-1 rocket plane. However, the Skystreak performed an important role in aeronautical research by flying for extended periods of time at transonic speeds, which freed the X-1 to fly for limited periods at supersonic speeds.