After publication of this work [1], we noted that we inadvertently failed to include the complete list of all coauthors. The full list of authors has now been added and the Authors’ contributions, Competing interests and Acknowledgment section have been modified accordingly.

Acknowledgements

This research has been supported by a Grant-in-Aid for Scientific Research (No.225803188) of the Japan Society for the Promotion of Science (JSPS). The authors thank Dr. DJ Skarzynski and Dr. MM Bah for the preparation of animals and for the assistance in CL collection.

HVV was supported by JSPS/Ronpaku dissertation program (No: VNM-11011). SHL was supported by a scholarship from the Ministry of Education, Culture, Sports, Science and Technology, Japan.

Competing interests

The authors declare that they have no competing interests.

Authors’ contributions

HVV participated in the in vitro experiments, data analysis and drafted the manuscript. SL carried out the in vitro experiments, data analysis and drafted the manuscript. ShY participated in data analysis and drafted the manuscript. HA carried out the in vitro experiments. KO participated in experimental design and discussion. TJA conceived of the study, participated in its design and performed in vivo and in vitro experiments. He also helped to draft the manuscript. All authors read and approved the final manuscript.

This article is published under license to BioMed Central Ltd. This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.