Koko’s capacity for language and empathy opened the minds and hearts of millions (Picture: Rohn Cohn/The Gorillia Foundation)

She was featured in multiple documentaries and appeared on the cover of National Geographic twice (Picture: Rohn Cohn/The Gorillia Foundation)

The foundation said in their statement: ‘Ginny’s impact has been profound and what she has taught us about the emotional capacity of gorillas and their cognitive abilities will continue to shape the world.

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‘The foundation will continue to honor Koko’s legacy and advance our mission with ongoing projects including conservation efforts in Africa, the great ape sanctuary on Maui, and a sign language application featuring Koko for the benefit of both gorillas and children.’