This is a joint project of Washington University, University of Vermont and the University of California, San Diego, partially supported by the National Science Foundation under a collaborative grant from the NSF-IBIV program ( DEB-1053566, DEB-1053554, DEB-1053559). Any opinions, findings, and conclusions or recommendations expressed in this material are those of the author(s) and do not necessarily reflect the views of the National Science Foundation. ·