var. spurium (トゲナシヤエムグラ), the type, western Eurasia (exotic to Japan, including Hokkaido)

索引

Galium spurium

var. echinospermon

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[1] near JR Otsu Station, Shiga Prefecture, on March 18 2012. [2/3] in the campus of Wuhan University (武漢大学), Hubei Province, China, on March 28 2012. [4] around Aoba Castle, northern Honshu, on March 22 2016.