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A question about bee orientation

The hypothetical question: Do bees orient more about North-South, arrangement of landmarks, or left-right?

The real life situation: I had a group of eight nucs that I needed to move about 50 yards while rotating about 180 degrees. I had a few choices. 1)Keep them in the same order, left to right. This choice would place the southern box in the northern position, and so forth. 2)I could move the nucs directly across the space, which would reverse the order, but keep the northern one the furthest north, and so forth. 3)About the arrangement of landmarks, I tried to keep the things around the nucs in the same relationship to them so that the bees would have as many visual cues as possible to prevent drifting.