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Denton and Hamilton were ever-presents during the Six Nations, while Jones only missed the final match in Italy due to concussion, and their absences are blows for a Scotland side seeking to arrest their worst run since 1998.

Scotland go into the Test with Australia on June 5 seeking to end a seven-match winless sequence, before playing the NSW Waratahs on June 10 and Tests against Fiji on June 16 and Samoa a week later.

Squad

Head coach Andy Robinson is poised to name his squad on May 16.

Jones' absence can be compensated for by the inclusion of Edinburgh's flying Dutch winger Tim Visser, who come June 12 will be eligible for Scotland on the grounds of three years' residency.

Missing Denton will stretch Scotland's back-row resources, with Saracens' Kelly Brown already absent after he dislocated his fibula in January, while Hamilton's indiscretion means he returns behind World Cup captain Alastair Kellock in the second row order.

Scotland team doctor James Robson said: "Lee suffered the fracture in the Heineken Cup semi-final loss to Ulster last month.

"Injuries of this nature typically resolve over the course of six to eight weeks, so he will not be available for the tour.

"David sustained the ankle injury in the final game of the league season against Treviso last weekend and subsequent scanning of the joint revealed damage to the ligaments.

"Following consultation with John McKinley, the ankle specialist at Spire Murrayfield Hospital, David will not require surgery but will now embark on a prolonged period of rehabilitation which will rule him out of the summer tour.