Mid-19th century England... the tramways that first appeared to transport coal have since been upgraded to railways. Numerous different companies using different railway gauges connected to villages, towns and cities across the country. Eventually the small companies were merged into the Big Four.

This expansion has two sets of rules. The first set of rules follows the gameplay of the Railways of the World series. The second set of rules is a new game from Martin Wallace that features a new share system and economic engine that builds upon the Railways of the World engine.

Railways of England and Wales debuted at Origins 2009. Now out of print, it was replaced in 2013 by Railways of Great Britain.