AUSTIN — Many of those who stayed booed. Many of those who played figured they’d soon be subjected to worse. And in the locker room after the latest unfathomable debacle at Royal-Memorial Stadium, Mack Brown gave it to them.

At Texas, where a program accustomed to competing for championships was inexplicably and humiliatingly dominated at home by a huge underdog for the second time this season, the stages of denial and disappointment officially ended Saturday.

After a jaw-dropping 28-21 loss to Iowa State, the Longhorns moved full-bore into anger.

They seethed during the traditional postgame playing of “The Eyes of Texas,” with defensive end Eddie Jones’ wild gesticulations looking more intimidating than anything UT had done in the four hours before. Then an infuriated Brown unleashed the kind of harsh, unfiltered emotion he’d never shown after even his most bitter previous defeats.

“What we did was totally unacceptable,” Brown said. “There’s no way to hide from that.”

More than once, Brown said his team “stunk.” More than once, he criticized his team’s “arrogance.” After conceding that Iowa State — which was blown out 52-0 by Oklahoma last week and had never defeated UT — was better coached than the Longhorns, Brown said he’s “mystified” by why his team wasn’t ready to play.

Iowa State (4-4, 2-2 in the Big 12) began the day ranked 112th nationally in total defense and 104th in total offense. The Cyclones had been outscored 120-27 in its last two games, and hadn’t won a game against a ranked opponent on the road in 20 years.

Texas (4-3, 2-2) was coming off what appeared to be a season-saving victory at then-No. 5 Nebraska. But despite the fact that they followed an impressive road performance at Texas Tech with an embarrassing home loss to UCLA in September, the Longhorns thought it was inconceivable it could happen again.

“Nobody for a second thought we could lose,” running back Tre’ Newton said.

That attitude, Brown said, was part of the problem. Although the game was officially a sellout with an announced crowd of 100,142, many of those seats remained empty at the time of the 11 a.m. kickoff, and the player’s disinterest seemed to match that of the fans.

Said Brown, who suffered back-to-back home losses for the first time in his 13 years as UT’s head coach: “I was worried in pregame, because they were standing out there with an arrogance.”

They did little to back that attitude up. With another sloppy start by an ineffective Texas offense and an uninspired Longhorns’ defensive effort, the Cyclones jumped out to a halftime 14-3 lead. UT committed four turnovers — the costliest coming when quarterback Garrett Gilbert was sacked and fumbled one play after the Longhorns intercepted a pass trailing 21-6 early in the fourth quarter.

“We came to play,” Iowa State coach Paul Rhodes said. “We were focused, resilient and we executed.”

In other words, they were the opposite of Texas. Gilbert threw for 344 yards and two touchdowns but threw three interceptions. The defense allowed Iowa State’s Alexander Robinson gash his way to 120 yards and two scores and put up little resistance on two key second-half drives.