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They also used their equations to estimate body masses of extinct carnivores. They predicted a body mass of 630 kg for an amphicyonid CA. major) that is also examined in the present study. This estimate will be commented on in the discussion. In general, dentally based estimates that include the widths of teeth have been suggested to be more dependent on diet than body mass (Damuth 1990, Fortelius 1990, Janis 1990). Anyonge (1993) made a thorough study of relationships between cross-sectional properties, distal articular surface area, lengths and circumferences of proximal limb bones (femur and humerus) and body mass in Felidae, Canidae, Ursidae, and Hyaenidae.

Leo) on the basis of original data and data from Nowak (1991). An exception is the lower carnassial, which is much shorter in leopards. The body mass range given for leopards is 28 to 90 kg, and for female lions 120182 kg. Body mass estimates for the amphicyonid based on head-body length lie within the leopard range. All the estimates based on skull length are higher; 117 to 393 kg. Occiput to orbit length gives estimates from 78 to 133 kg. The estimates based on the lower carnassial vary between 29 and 218 kg, being highest when the felid equation is used and lowest when the canid equation is used (Table 3).

The distribution of the species in the plot is in accordance with their placement in the feeding categories above. The omnivorous P. bavaricus has a short blade characteristic of a less carnivorous mode of life. The bone-crushing mesocarni- 0 P. styriacus [J P. bavaricus • C. helbingi b T. bourgeoisi vores have longer blades than P. bavaricus, but shorter' than the species that were classified as hypercarnivores. The blade is shorter in A. major than in P. steinheimensis or P. styriacus. C. helbingi, which was classified as a mesocarnivore, has a blade length similar to that of P.