1Center for Research on Genomics and Global Health, National Human Genome Research Institute, National Institutes of Health, Bethesda, MD 20852, USA2Department of Medicine, University of Ibadan, Ibadan 200284, Nigeria3Department of Medicine, University of Lagos, Lagos 101017, Nigeria4Department of Medicine, University of Nigeria Teaching Hospital, Enugu 400001, Nigeria5Department of Medicine and Therapeutics, University of Ghana Medical School, Accra, Ghana6Department of Medicine, University of Science and Technology, Kumasi, Ghana7Institute of Human Virology, School of Medicine, University of Maryland, Baltimore, MD 21201, USA