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All the animals were rounded up within half an hour, and police said the dogs were handed over to someone in the local area to look after while investigations go on and the owners are tracked down.

A police spokesman said: “Inquiries continue into the dogs being loose on the A55.

“But police can confirm the fox died after being struck by a vehicle.”

Speaking after the incident, Anglesey’s Chief Inspector Mark Armstrong said: “Local and Roads Policing Officers were quickly on the scene and fortunately the dogs’ presence close to the A55 caused no collisions and within 30 minutes of the call the dogs were recovered and handed over to a local person.

“I’m grateful to the member of the public who made the initial call as this may well have prevented a more serious incident.”