1Department for the Monitoring and Protection of the Environment and for Biodiversity Conservation, Italian National Institute for Environmental Protection and Research (ISPRA), via Vitaliano Brancati 60, I-00144 Roma, Italy.

2Centre for the Conservation of Biodiversity (CCB), Department of Environment and Life Science (DISVA), University of Cagliari, v.le Sant'Ignazio da Laconi 11-13, I-09123 Cagliari, Italy.