The parasites first three larval stages could be obtained and studied by the infestation of Orthoptera, Dermaptera and Dictyoptera. Its evolution in the definitive host has been observed in two Rodents Muridae: Apodemus sylvaticus L. and Mus musculus L. and in a Cricetidae: Calomys callosus (Rengger). During the larval development, the pharyngial structures as well as the teeth and lips are altered.

At the end of the second larval stage they re call those of the Gongylonematinae.

At the third stage they are comparable to those of the infesting larvae of Spirocercinae and Ascaropsinae.

At the fourth stage they are similar to those of the adults of some Ascaropsinae : Pygarginema and Physocephalus.

The adult of Mastophorus muris which possesses separated and differentiated labial lobes is an evolutive form.

The family Spiruridae is reclassified according to the analysis of its larval forms and the host lists of its different subfamilies.

The new subfamily Mastophorinae n. subfam. is retrieved from the Spirurinae on account of its being derived from the Ascaropsinae; the new subfamily Hartertiinae n. subfam. (including the genus Hartertia, Seurat, 1914 and Alainchabaudia Mawson, 1968) is separated from the other Spiruridae subfamilies on the grounds of having a different cephalic morphogenesis.

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