Birds (class Aves or clade Avialae) are feathered, winged, bipedal, endothermic (warm-blooded), egg-laying, vertebrate animals. With around 10,000 living species, they are the most speciose class of tetrapod vertebrates. All present species belong to the subclass Neornithes, and inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from the 5 cm (2 in) Bee Hummingbird to the 2.75 m (9 ft) Ostrich. The fossil record indicates that birds emerged within theropod dinosaurs during the Jurassic period, around 150 million years ago. Birds are the only members of the clade originating with the earliest dinosaurs to have survived the Cretaceous–Paleogene extinction event 66 million years ago.Modern birds are characterised by feathers, a beak with no teeth, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a lightweight but strong skeleton. All living species of birds have wings; the most recent species without wings was the moa, which is generally considered to have become extinct in the 16th century. Wings are evolved forelimbs, and most bird species can fly. Flightless birds include ratites, penguins, and a number of diverse endemic island species. Some species of birds, particularly penguins and members of the Anatidae family, are adapted to swim. Birds also have unique digestive and respiratory systems that are highly adapted for flight. Some birds, especially corvids and parrots, are among the most intelligent animal species; a number of bird species have been observed manufacturing and using tools, and many social species exhibit cultural transmission of knowledge across generations.