Brian Lima is one of five changes in Samoa's starting XV for their must-win match against England in Nantes.

Lima will partner Seilala Mapusua at inside centre but Samoa will be without key goal-kicker Gavin Williams, who has been ruled out with a broken finger.

Loki Crichton comes in at full-back for the injured Williams, while David Lemi moves to the right wing.

Number 8 Henry Tuilagi and fly-half Eliota Fuimaono-Sapolu also return, along with prop Kas Lealamanua.

Lima became the first player to appear at five World Cups when he came on as a replacement in Samoa's 59-7 defeat to South Africa.

However, the veteran Bristol centre, nicknamed 'the Chiropractor' for his crunching tackles, was knocked out minutes after coming on and was subsequently ruled out of his side's 19-15 loss to Tonga on Sunday.

Both sides need a victory to maintain their hopes of reaching the latter stages of the tournament, a fact not lost on Samoa coach Michael Jones.

"We're all very clear on the significance of this game and what it means in terms of what we represent in terms of our country, our families, all the good things we play for," he said.

"If we get it wrong this week - we don't even want to think about that. The boys are excited, they're very bubbly and very focused."