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Study of biology at TH Darmstadt; doctoral thesis on “Queen Polymorphism in the North American ant Leptothorax sp. A”, supervised by Prof. A. Buschinger and supported by German Academic Scholarship Foundation; doctoral examination in May 1988.

June 1988 to June 1989 post-doc in the lab of Prof. E.O. Wilson, Harvard University, Cambridge, Mass. Studies on the systematics, population biology, and ecology of North American ants of the genus Leptothorax; field work in Quebec, New England, Alberta and British Columbia.

September 1989 – Februar 1995 scientific assistant at the chair of Prof. Dr. B. Hölldobler, University of Würzburg; establishment of the research areas population biology and sociogenetics; studies on reproductive conflict and conflict resolution in ant societies. Field studies in North America, India, Brazil, Turkey.

July 1994 Habilitation in zoology with a thesis on "Cooperation and Conflict in Ant Societies",

March 1995 – September 1996 Heisenberg Fellowship

October 1996 – March 2000 Professor of Zoology at University of Erlangen-Nürnberg