Bergmans, W. 1979. Taxonomy and zoogeography of the fruit bats of the People's Republic of Congo with notes on their reproductive biology (Mammalia, Megachiroptera). Bijdragen tot de Dierkunde 48:161-186.

Bhatnagar, K. P., H. D. Frahm, and H. Stephan. 1990. The megachiropteran pineal organ: a comparative morphological and volumetric invesigation with special emphasis on the remarkably large pineal of Dobsonia praedatrix. Journal of Anat.