Madsen, Wageningen University and Research, Wageningen, Nederland
;
Crooijmans, Wageningen University and Research, Wageningen, Nederland
;
Schachtschneider, University of Illinois at Chicago, Chicago, United States of America
;
Groenen, Wageningen University and Research, Wageningen, Nederland

Ryan Morrison, University of Warwick, Coventry, United Kingdom
;
Robin Allaby, University of Warwick, Coventry, United Kingdom
;
Logan Kistler, Smithsonian National Museum of Natural History, Washington, United States of America