The car in front of his stopped for no reason. Another driver, not involved in what was about to unfold, stopped behind him. Boulis, a prominent South Florida businessman, had nowhere to go as a black Ford Mustang pulled up beside him.

The driver of the Mustang pulled out a gun. Boulis held out his hand when the first shot was fired. He could not have been more than two feet away from the weapon when it fired, according to the doctor who performed the autopsy.

Jurors in the murder trial of Anthony "Big Tony" Moscatiello and Anthony "Little Tony" Ferrari learned Tuesday exactly how Boulis — the founder of Miami Subs who was locked in a power struggle over the lucrative SunCruz Casino business he sold in a deal laced with fraud — met his demise on Feb. 6, 2001.

Eroston Price, at the time an assistant Broward Medical Examiner, said Boulis suffered four gunshot wounds. Two were minor, she said. The first grazed his forehead. The second, his left upper arm. The third shot entered his left upper back.

Boulis could have survived any of those wounds, but not the fourth, Price said. The last shot entered Boulis' back, went through his aorta and was retrieved from underneath his chin.

"This was the fatal wound," Price said.

The pattern of gunshot wounds indicates Boulis was turning away from his assailant as the shots were fired.

The medical testimony was paired with a firsthand account from the only living eyewitness to the shooting. Robert Puskarich, who happened to be driving southbound behind Boulis' BMW on Miami Road just east of Federal Highway in Fort Lauderdale, described the shooting and said he immediately feared for his own life.

"I went down in my seat, toward the passenger side," Puskarich said. "I was afraid he was going to shoot me."

When he raised his head, Puskarich said, the Mustang was still there and he was face to face with the driver — the man he had just seen shoot Boulis. But there were no more shots fired. The man calmly drove away, Puskarich said.

Prosecutors believe the gunman was a man named John Gurino, who was later killed in a shooting in Boca Raton. Gurino was hired, prosecutors say, by Moscatiello and Ferrari. They face the death penalty if convicted.

None of the testimony so far has tied either man to the crime. Prosecutors in the coming days expect to call Adam Kidan, the business rival who hired Moscatiello for protection, and James "Pudgy" Fiorillo, who was originally a co-defendant in the case and admitted to his role in a 2011 plea deal.