Background*

Dr. Joshua S. Boger, Ph.D., has been an Executive Director of Vertex Pharmaceuticals Incorporated since 1989. He has been a Director of ActivBiotics, Inc. since December 2005 and Massachusetts Life Sciences Center since August 2011. He serves as a Director of The Hastings Center, Inc. He serves as a Director of Biotechnology Industry Organization. He founded Vertex Pharmaceuticals Incorporated in 1989 and served as its Executive Chairman from 1997 to May 2006 and also ... its Chief Executive Officer since 1992 until May 23, 2009. Dr. Boger served as the President of Vertex Pharmaceuticals Incorporated from May 11, 2005 to February 2009 and from 1989 to December 2000 and as its Chief Scientific Officer from 1989 to May 1992. He served as Senior Director of Basic Chemistry of Merck Sharp & Dohme Research Laboratories in Rahway, New Jersey, where he headed both the Department of Medicinal Chemistry of Immunology & Inflammation and the Department of Biophysical Chemistry. He began his industry career at Merck in 1978. Dr. Boger served as Chairman of Healthcare of Biotechnology Industry Organization. He served as a Director of Metaphore Pharmaceuticals Inc. Dr. Boger holds a B.A. in Chemistry and Philosophy from Wesleyan University and M.S. and Ph.D. degrees in Chemistry from Harvard University.

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