Perennial forest-floor herbs, mostly with slender creeping stems which root at the nodes and have fibrous roots

Leaves

Leaves opposite, with mostly long petioles; blades ovate-cordate to rounded-reniform; stipules interpetiolar, ovate, entire or bilobed at the apex

Flowers

Flowers hermaphrodite, sometimes heterostylous, mostly in terminal umbels or sometimes solitary, often on long peduncles held erect from the main stems, occasionally with an involucre of quite conspicuous bracts

Flowers hermaphrodite, sometimes heterostylous, mostly in terminal umbels or sometimes solitary, often on long peduncles held erect from the main stems, occasionally with an involucre of quite conspicuous bracts.

Delprete, P.G. & Cortés-B., R. (2006 publ. 2007). A synopsis of the Rubiaceae of the states of Mato Grosso and Mato Grosso do Sul, Brazil, with a key to genera, and a preliminary species list Revista de Biologia Neotropical 3: 13-96.

Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.